422 DR. M. WATSON ON THE MALE [Apr. 16 ( 



tion of these glands in the latter sex I pointed out that in respect of 

 them H. crocuta agrees more closely with Proteles than with any 

 species of its own genus, and may merely add that the present dis- 

 section has enabled me to confirm Dr. Murie's observation l that in 

 the male animal as in the female an anal-gland pouch is present. 

 Mr. Busk a had previously convinced himself " that no trace of a 

 pouch between the root of the tail and the anus exists, at any rate 

 in the male of this species ;" and Dr. Murie, in deference to Prof. 

 Kaup's 3 attempt to classify the Hyaenas in accordance with the 

 presence or absence of this pouch, was not prepared to deny that 

 it might occasionally be absent in the male. I think this matter 

 may now be considered settled, and may merely observe that Gold- 

 smith 4 appears to have been aware of the presence of this pouch in 

 every species of Hyaena before any of the authors referred to had 

 written on the subject. 



With reference to the male organs of allied species, I find that 

 Daubenton 5 describes and figures those of H. striata 5 and these are 

 also referred to by John Hunter 6 and Owen T . Cuvier 8 gives an 

 account of the male organs of Hyeena without indicating the species 

 to which they belong ; but from the agreement between his observa- 

 tions and those of the authors just named, I presume that H. striata 

 formed the subject of his investigations. Of the male organs of H. 

 brunnea, so far as I cau ascertain, we know nothing ; but of those 

 of the closely allied genus Proteles, Prof. Flower 9 gives an accu- 

 rate description. H. striata and Proteles agree with H. crocuta 

 with respect to the form and position of the scrotum, the form of 

 the testicle, and the absence of vesiculae seminales, whilst they differ 

 from the last-named species in the possession of a prostate gland. 

 Mr. Flower, it is true, refers to the presence of a prostate gland in 

 H. crocuta 10 ; but of the absence of this in the specimen I examined 

 I was careful to convince myself; and until further observations are 

 made, it is impossible to reconcile these opposing statements regard- 

 ing a matter of fact. In H. striata the prostate is large and con- 

 sists of two kidney-shaped lobes, whilst in Proteles it consists of a 

 single mass of large size having " the appearance of a bilobed disk." 

 Cowper's glands are large and of the same form in all three species, 

 which likewise agree in the form of the bladder and the very mus- 

 cular character of the ureters. With respect to the vesicula pros- 

 tatica, Leuckart u describes and figures it in H. striata as a minute 

 flask-shaped cul-de-sac projecting beyond the urethral wall and 

 lying in the interval between the lobes of the prostate. He s&ys, 

 moreover, that in that animal it does not open into the urethra. 



I Trans. Zool. Soc. vii. p. 503. 



a Journ. Proc. Linn. Soc. vol. ix. p. 71. ' Isis, 1828, p. 1144. 



* Animated Nature, " Hysena." 



* Buffon's Hist. Nat. torn. ix. 



6 ' Essays and Observations,' by Owen, vol. ii. p. 58. 



' ' Anatomy of Vertebrates,' vol. iii. p. 671. 



8 ' Anatomie Compare^,' vol. v. 



» Proc. Zool. Soc. 1869, p. 491. 10 Ibid. p. 493. 



II ' Cyclopaedia of Anatomy,' vol. iv. Art. "Vesicula prostatica." 



