1878.] GENERATIVE ORGANS OK HY^NA CROCUTA. -J 23 



Mr. Flower found no distinct uterus masculinus in Proteles ■ but as 



could S? medmn • Idge ° n thC fl0 ° r ° f the Urethra ' S which 1 

 could detect a very minute aperture, the arrangement appears to be 



very similar to that described above in H. crocuta. Tn re pect 



therefore of the form of the vesicula prostatic, //. J/! 



more closely with Proteles than with k striata, differing from the 



w 1 and"n U tt a f 'rtltt ^ ^ ^ Ct b ^° nd the **** 

 sarnP A ? ?*' , U commuD ^ates with the canal of the 



tZ t.h™° rd / f . LeUCka - t l * iS aU extreme] y rare occurrence 

 ? s i 7 ? ^ reUtl - ° pen lnt0 th e vesicula prostatica; but this 

 is certainly he case in H. crocuta. The penis of the Spotted 



onHo t 7 reSemb , leS V hat ° f H - StHata ™ d of steles, dlffe hg 

 only from these in the absence of a conical bodv of car tilaginouf 

 cons stance which has been described by Cuvier in the glansCn" 

 o the former and by Professor Flower in that of the latter. P The 

 glans penis in all of them is invested by small recurved spines. 



Comparison of the Male and Female Organs of H. crocuta - 

 When describing the female organs, I remarked that "the a range- 

 rnent of tl „ h ag ^ & re8emblance h * 



ot the males in o her species," and that, if we supposed the vesicula 



EuTtl th l ktter ^ larged t0 the Slze ° f the female ute - w 

 a rale'" n, 'f b r UCe °i " F ° State gla " d ' haVe an almost identical 

 S ! T'f^ 7 paSSageS ° f b0th sexes ' The abse »ce, 



make f t n ' ° f ? e Pr ° State S land in tbe male H - vocuta 



make, the resemblance between the male and female organs of this 

 specie, even greater than I anticipated. And here I may be pe - 

 mi ted to observe that nowhere in the group of mammals is the tmth 



tl e ho™ « I S WlUC1 em Wologists have arrived respecting 

 the homologies of the various parts of the sexual apparatus in the 

 MS ^V* beautlfu % sh °wn as in the animal underconsideration 



thl y i U u remain re f rdin g the similarity of plan upon which 

 these are built un. it wnn hi ho Q t ™™ ,i:„^„h/j u.. _ • ^ *»»^u 



f , _ " , .,; "&'»"»'s me smniaruy or plan upon which 



these are built up, ,t would be at once dispelled by an inspection of 

 he sexual organs of Byana crocuta. In both sexes thereha short 

 urethra opening close to the mouth of the uterus, which organ in 

 accordance with functional requirements, is of large size in the female 

 but is reduced to a minimum in the male. In both there is a urino- 

 gemal canal extending from the junction of the urinary and sexual 

 canals to the extremity of the penis or clitoris, according to sex ; and 

 in both there are two Cowperian glands of large size opening into 

 tha t canal close to the root of the penis or clitoris. A complriZ 



tw S, I 6 ™ 1 gemtal ,°T nS ° f the two sexes P roves conclusively 

 that the lower part of the so-called prostatic, together with the 

 membranous portion of the male urethra, are homologous with 

 the commencement of the urino-genital canal of the female, which 

 canal, in the majority of mammals, is so metamorphosed to form 

 the vestibule, that its homologue in the other sex is by no means 

 readily recognizable. This homology is further substantiated, in 

 the animal we are considering, by the size and form of Cowper's 

 glands, and the points of entrance of the ducts of these into the 

 urino-genital canal in each sex. Lastly, the homology of the so- 



