6/8 DR. WATSON ON DASYPUS SEXCINCTUS. [JlUie 18, 



by the junction of the two crura alone, and possesses neither bulb 

 nor glans. With regard to the other muscles of the penis, these 

 closely resemble those already described in Chlamydophorus, with 

 this difference, that the single mass formed by the bulbo-cavernosi 

 muscles in the latter are here represented by two distinct and 

 powerful fleshy bands which envelop the under surface of each 

 cms previous to its junction with its fellow. The testicle differs 

 in position in Dasypus from that which it occupies in Chlamydo- 

 phorus, being attached to the inguinal region of the anterior ab- 

 dominal wall in the former, whilst in the latter it lies behind the 

 kidney. The globus major in Dasypus is closely applied to the 

 anterior extremity of the testicle, and does not form the loop- 

 like arrangement seen in Chlamydophorus. The vasa deferentia in 

 both are non-convoluted ; nor is there any trace of vesiculse semi- 

 nales in either. In both, a prostate and Cowper's glands are 

 present, the former being traversed by the vasa deferentia in their 

 course to the urethra, whilst the latter, although occupying the same 

 position, are relatively considerably smaller in Dasypus than in Chla- 

 mydophorus. The bladder and urethra are similar in both genera, 

 the only difference being that the walls of the intrapelvic portion of 

 the latter are much more muscular in Chlamydophorus than in Da- 

 sypus, and that in the latter a well-marked caput gallinaginis, of a 

 pyramidal form, is present, which is not represented in the former. 

 In neither is there any trace of a vesicula prostatica. 



It will thus be seen that in respect of the male sexual organs the 

 resemblance between Chlamydophorus and Dasypus sexcinctus is very 

 striking — a resemblance which, in respect of other organs, has been 

 already pointed out by Prof Garrod '. As shown by that anato- 

 mist, Chlamydophorus agrees also with Tolypeutes and Xenurus in 

 the absence of vesiculae seminales and the simple condition of the 

 glans penis, in both of which respects it differs from Tatusia. The 

 presence of Cowper's glands constitutes a further element of agree- 

 ment between Chlamydophorus on the one hand, and Dasypus and 

 Xenurus on the other. These glands are also present, according to 

 Dr. Murie 2 , in Tolypeutes ; but whether in Tatusia I cannot say, as 

 with regard to the latter we possess no precise information. So far, 

 then, as the observations above recorded go, they tend to strengthen 

 the conclusions already arrived at by Mr. Garrod upon other grounds 

 with regard to the close relationship which exists between Chlamydo- 

 phorus and Dasypus. Professor Macalister, as the result of his 

 elaborate investigation into the muscular anatomy of the Edentata, 

 arrives at the conclusion that " the position of Chlamydophorus will 



be seen to be plainly among the Dasypodidse, and very close 



to Tatusia." The arrangement, however, of the sexual organs, 

 including the nipples, together with that of the alimentary canal, 

 which in Dasypus and Chlamydophorus is provided with two colic 

 caeca, these last being absent in the other genera above referred 

 to 3 , appear rather to point to a closer relationship between Chla- 



1 P. Z. S. 1878, p. 222. s Trans. Linn. Boo. vol. xxx. , 



3 See Prof. Garrod's paper. 



