684 MESSRS. GARROD AND TURNER ON [JunelS, 



membrane consisted of a gland layer and a crypt layer. The gland 

 layer was next the muscular coat, and consisted of elongated tubular 

 glands, somewhat tortuous and occasionally bifurcating. In the 

 vertical sections the glands were cut across so that the tubes were 

 sometimes transversely, at others obliquely, at others longitudinally 

 divided, and here and there the stem of a gland could be seen pass- 

 ing obliquely through the crypt-layer to open on the surface in the 

 manner already described. The glands were lined by a columnar 

 epithelium, and possessed a central lumen. The glands were neither 

 so numerous nor so distinct, neither did they bifurcate so frequently 

 as do the utricular glands in the Pig and the Cetacea. 



The crypt layer contained the numerous depressions already re- 

 ferred to for the lodgment of the villi of the chorion. The epithe- 

 lium lining the crypts had, as a rule, disappeared ; so that it was only 

 in exceptional localities that it could be seen in situ, where it ap- 

 peared to consist of cells, the type form of which was columnar, 

 though modifications of that shape occurred. The subepithelial 

 connective tissue contained a large proportion of corpuscles, some of 

 which were fusiform, others polygonal, others of the rounded form 

 of white blood-corpuscles. This tissue was more compact where it 

 formed the walls of the crypts ; but deeper in the mucosa, as it ap- 

 proached the glandular layer and the muscular coat, it had an areo- 

 lated character. The vessels of the uterus were not injected ; but 

 there can be no doubt that, if they had been so, the walls of the 

 crypts would have been seen to contain an abundant freely anasto- 

 mosing network of capillaries, such as exist in the corresponding 

 crypts in the Cetacea, the Mare, the Pig, and the Lemurs. In 

 sections through the wall of the uterus, that had been stained with 

 hsematoxylin, a well-defined band, coloured with the blue pigmeut, 

 marked the junction of the deep surface of the mucous membrane 

 and its glands with the muscular coat. This band in all probability 

 was the muscularis mucosae. In Hyomoschus, as in other animals 

 possessing a diffused placenta, the uterine glands have no relation, as 

 regards numbers or termination, to the crypts. The crypts are in- 

 finitely more numerous than the glands, and are not to be regarded 

 as formed by a dilatation of their mouths, but are new formations 

 during pregnancy, due to hypertrophy and folding of the mucous 

 membrane so as closely to adapt it to the irregular villous surface 

 of the foetal chorion. 



The chorion extended from the tip of the left uterine cornu, through 

 the corpus uteri, to the tip of the right uterine cornu. The left horn 

 of the chorion, which contained the foetus, was longer and much 

 more capacious than the right horn. The tip of each horn of the 

 chorion was in close relation to the orifice of each Fallopian tube ; and 

 close to the tip the free surface of the chorion was over a very limited 

 area smooth and non-villous. That part of the chorion situated in 

 the corpus uteri, immediately opposite the os uteri, presented a cir- 

 cular non-villous surface about the size of a shilling. This surface, 

 though without villi, was folded so as to adapt it to the correspond- 

 ing folds of the uterine mucosa in the same locality. A portion of 



