1 8 SCOTT. [Vol. V. 



modern types. But even the facial portion of the maxillaries is 

 very low, especially in front, and the nasal chamber is not nearly 

 so high as in the camel and llama. Just below the line of 

 suture with the nasals there is a long depression in the maxilla- 

 ries, which is constant in all the specimens, and cannot therefore 

 be entirely due to pressure. Seen in section the nasal passage 

 thus has a curious shape, very narrow above, but broad and very 

 low beneath. Posteriorly the maxillary sends up a short process 

 between the lachrymal and nasal, which reaches the frontal, as 

 in Camelus, while in Anchenia the vacuity developed at this point 

 prevents more than the minutest contact with the frontal. The 

 alveolar border is prolonged much farther back of the last molar 

 than in the existing forms. The face of Pcebrotherium does not 

 exhibit the sudden constriction in advance of pm. 3, which is so 

 characteristic of recent Tylopoda, but tapers gradually to the 

 muzzle, and as the premolars are still present in unreduced 

 number, there is a much shorter edentulous region of the maxil- 

 lary than in the camel and llama. 



The premaxillaries are only partially preserved in any speci- 

 men which I have examined, but it can be seen that the ascend- 

 ing rami are short and nearly vertical, and that they are in 

 contact with the nasals. The anterior nares have thus the low, 

 narrow shape and nearly vertical position which occurs in Ano- 

 plotlierium, Cainotherium, Oreodon, and in fact most of the 

 primitive selenodonts, and very different from the long, obliquely 

 placed opening of the recent tylopodans. Apparently the pala- 

 tine plates of the. premaxillaries are large in proportion and but 

 little reduced by the incisive foramina. The palatine processes 

 of the maxillaries are long, narrow, and tapering anteriorly, 

 giving to the palate the triangular shape, very narrow in front, 

 which is characteristic of the Camelidce, but there is hardly any 

 indication of the constriction of the palatal surface at the 

 diastema behind the first premolar and re-expansion at the canine 

 alveoli, which is so marked in the recent genera. The palatine 

 fissure between the alveolus and the posterior nares is very 

 shallow, though subject to considerable variation : in some speci- 

 mens it hardly reaches the third molar. 



The palatines are narrow and are intermediate in character 

 between those of Camelus and those of Auchenia ; they extend 

 well forward, as far as the first molar, and have their anterior 



