94 BOTANICAL GAZETTE | AUGUST 
of attraction is exhibited in A. candida during mitosis, It often 
results in a spindle nearly twice the normal length and propor- 
tionately narrower. Such cases are represented in figs. 15-20. 
The consequence of mitosis under these conditions is that several 
daughter nuclei remain anchored to the coenocentrum ( figs. 77, 
ats 22). Whether there is an organic attachment or merely an 
imbedding of a projection is impossible to say, but certainly 
these daughter nuclei protrude a long pseudopodium-like exten- 
sion which penetrates the coenocentrum to a considerable depth. 
In late stages, as the nuclei pass to the periplasm, fewer are 
found attached to the coenocentrum, with the result that eventu- 
ally only one remains. This nucleus enlarges greatly, and is 
often found lying in the cytoplasm in such a position as to 
suggest that it had been fixed while swaying to and fro on the 
stalk-like pseudopodium which attaches it to the coenocentrum. 
The migration and attachment of the nuclei to the coeno- 
centrum seem inexplicable on any basis save that of chemo- 
tactic influence. 
The growth of the nucleus while in contact with this struc- 
ture, both in this species and in A. Tragopogonis, gives evidence 
that the coenocentrum functions as an organ of nutrition for the 
one surviving nucleus. In some respects the coenocentrum 
resembles a trophoplast, but the presence of granules, probably 
of kinoplasmic nature, renders the analogy less complete. It 
seems also to arise de novo in each oogonium, a character not in 
agreement with the theories most generally accepted. The 
evidence that such structures always arise from preexisting ones 
is not conclusive, however, and the work of Davis (1899) on 
Anthoceros may indicate that the chloroplast in that plant 
arises dé novo in each sporophyte from the cytoplasm of the 
spore mother cells, although the author does not definitely draw 
that conclusion. If such be the case, the similarity between the 
coenocentrum and the chloroplast is closer, Evidence of high 
specialization in chloroplasts is given, moreover, in observations 
of Oltmanns on Coleochaete (1898), and of Davis on Anthoceros 
(1899), where the chloroplasts divide in advance of the nuclei, so 
