1901] EREMOSPHAERA AND EXCENTROSPHAERA 317 
a more complicated life history, including the formation of 
zoospores, was suspected. Henfrey, in his discussion of the 
plant already referred to, suggested the possibility of anther- 
ozoids, but this condition was undoubtedly due to the presence 
of some parasite, which Henfrey himself recognized as a possible 
cause, 
While considerable doubt has been cast upon Chodat’s inter- 
pretation of what he saw, the fact that certain of the stages 
which he describes certainly exist in material found in this 
country, makes it necessary that a most careful search be made 
for all the forms supposed by him to be connected with Ere- 
mosphaera. In addition to the well-known division into two or 
four spheres, Chodat (1) describes a kind of sporangium-building 
condition where division goes on until the diameter of the ulti- 
mate cells is ten to twenty times smaller than at first. This 
stage I have been unable to find, and nothing of the sort seems 
to occur in the material I studied, either in gross or pure 
cultures. In addition to these non-motile spores, Chodat des- 
cribes ciliated spores. Just how and when these may be formed 
is not clear, but it is to be inferred that they arise from some 
‘“‘palmella” condition and not directly from the adult plant. 
These zoospores, according to Chodat, are usually elliptical, and 
are always surrounded by a gelatinous coat, reminding one of 
certain forms of Chlamydomonas. Each zoospore has a red eye 
Spot, a radiating chromatophore, and an evident nucleus. In 
most cases there are two cilia, but exceptionally three. The 
question as to whether they actually had four remained unde- 
cided, as did also the possibility of their conjugation. They 
seem to have been irregular both as to size and shape, for Chodat 
(1) says, ‘da ihre Mutterzellen an Grosse sehr verschieden sein 
konnen, so sind infolgedessen die Zoosporen auch sehr verschie- 
den, sowohl was die Grosse als auch die Form anbetrifft.” 
Although a most careful search has been made among mate- 
rial growing under all conditions, not the slightest trace has ever 
been discovered of any motile spore in evident relation to Ere- 
mosphaera. Occasionally plants would be found of a lighter 
