﻿2 
  BOTANICAL 
  GAZETTE, 
  [January, 
  

  

  soon 
  as 
  w 
  

  

  tosporangiate 
  

  

  A 
  1 
  ■ 
  * 
  

  

  ^^e 
  attempt 
  to 
  homologize 
  the 
  sporophyte 
  of 
  any 
  lep- 
  

   pate 
  fern 
  with 
  that 
  of 
  any 
  known 
  alga 
  or 
  brvophvte. 
  

  

  of 
  

   d 
  

  

  Anything 
  in 
  the 
  least 
  resembling 
  the 
  epidermal 
  sporangia 
  c 
  

   the 
  Filices 
  is 
  absolutely 
  unknown 
  outside 
  of 
  the 
  group, 
  ana 
  

   the 
  high 
  degree 
  of 
  development 
  of 
  the 
  body 
  of 
  even 
  the 
  sim- 
  

   plest 
  of 
  these 
  is 
  separated 
  by 
  an 
  immense 
  distance 
  from 
  the 
  

   sporanguim 
  of 
  any 
  known 
  form 
  among 
  either 
  bryophytes 
  or 
  

   algae. 
  It 
  must 
  be 
  borne 
  in 
  mind 
  that 
  the 
  prothallium 
  of 
  a 
  

   Hymenophyllum 
  corresponds 
  not 
  merely 
  to 
  the 
  protonema 
  of 
  

   a 
  moss, 
  but 
  to 
  the 
  protonema 
  //?« 
  the 
  leafy 
  sexual 
  plant. 
  

  

  If 
  we 
  go 
  back 
  to 
  the 
  older 
  botanists 
  we 
  find 
  a 
  different 
  

   view 
  as 
  to 
  the 
  origin 
  of 
  the 
  pteridophytes, 
  tiiough 
  here 
  also 
  

   the 
  leptosporangiate 
  ferns 
  seem 
  to 
  be" 
  regarded 
  as 
  the 
  more 
  

   primitive 
  forms. 
  According 
  to 
  this 
  earlier 
  view 
  the 
  pterido- 
  

   phytes 
  probably 
  originated 
  from 
  some 
  simple 
  form 
  allied 
  to 
  

   the 
  Hepaticae. 
  The 
  evident 
  resemblance 
  between 
  sucii 
  a 
  

   form 
  as 
  Anthoceros, 
  for 
  example, 
  and 
  an 
  ordinarv 
  fern-pro- 
  

   thallium 
  IS 
  obvious, 
  and 
  I 
  hope 
  to 
  show 
  that 
  there 
  is 
  some 
  

   good 
  reason 
  for 
  reconsidering, 
  at 
  least, 
  this 
  older 
  view. 
  

  

  L.eitO"pV|2 
  pollc 
  affat-.f;^^ 
  4-^ 
  tU„ 
  11 
  1 
  »...,-. 
  

  

  _. 
  - 
  j^, 
  "V 
  ^v-c^ou, 
  lino 
  ijiut.1 
  view. 
  

  

  l^eitgeb^ 
  calls 
  attention 
  to 
  the 
  well-known 
  but 
  significant 
  

   t 
  that 
  among 
  the 
  Anthocerote^e 
  alone, 
  of 
  all 
  known 
  bry- 
  

   ophytes, 
  the 
  growth 
  of 
  the 
  sporogonium 
  is 
  unlimited, 
  coii- 
  

   tmuing 
  to 
  grow 
  at 
  the 
  base 
  as 
  long 
  as 
  the 
  plant 
  lives. 
  The 
  

  

  Sr/rT 
  /' 
  ''^'''' 
  -f 
  ^?''^^^ 
  1^^^^ 
  and 
  contains 
  abundant 
  

   cl7nJ^Tl7'^\ 
  '"''} 
  V"'^' 
  intercellular 
  spaces 
  communi- 
  

   cating 
  with 
  stomata 
  of 
  the 
  same 
  structure 
  as 
  those 
  of 
  the 
  

  

  c 
  fL'd 
  Ti?n"t'''".''' 
  '^i^^^^ 
  'Z 
  ^^^ 
  '' 
  assimilation 
  is 
  con 
  

   ceined, 
  it 
  is 
  quite 
  independent 
  of 
  the 
  oophvte. 
  Add 
  to 
  this 
  

  

  that 
  t^ie 
  columella, 
  both 
  in 
  structure 
  and 
  porsition 
  c 
  oselv 
  re 
  

   sembles 
  the 
  young 
  axial 
  fibro-vascular 
  bundles^f 
  the 
  em' 
  

   "[^of 
  Chn^P'^"'^' 
  ^"' 
  r 
  ''' 
  ^-^^ 
  closel 
  the 
  sporogo: 
  

  

  to 
  show, 
  is 
  probablv 
  the 
  nfr^-f-' 
  V.^' 
  ^^ 
  ^ 
  ^^^^^1 
  endeavor 
  

  

  cine*. 
  ^ 
  ^ 
  ^^^ 
  primitive 
  condition 
  among 
  the 
  Fili- 
  

  

  