594 ALLIS. [Vor. XII. 
branches of the facialis. From its base the anterior root of 
the ganglion arises. It isa short, stout root, running inward 
and backward to the anterior and lateral corner of the medulla, 
where it has its origin. It is here distinctly double, having a 
_dorsal posterior portion and a ventral anterior one. The former 
arises by fibres that spread and extend backward in the brain, 
parallel to the outer surface of the brain, almost to the level of 
the origin of the facialis. The other, or ventral portion, arises 
in the brain as two bundles, which extend, as bundles, backward 
and medianward into the brain. The anterior bundle arises, as 
seen in horizontal sections, in front of the ventricular cavity of 
the brain, from the anterior, upturned end of the tract from 
which the ventral motor component of the facialis arises, and 
which I take to be the posterior longitudinal fasciculus of 
Osborne in cryptobranchs. This bundle is the motor V, or V 
minor, of Strong in Rana. 
The fibres of the antero-ventral part of the anterior root of 
the main ganglion form the larger part, if not all, of the anterior 
of the two bundles of fibres that traverse the ganglion. From 
this transverse bundle the truncus maxillaris in large part arises, 
the bundle turning downward in that part of the ganglion that 
lies external to the orbital vein, and then outward and forward 
into the truncus. In one 20 mm. larva the transverse bundle, 
or commissure, was distinctly double, the fibres of both parts 
arising from the antero-ventral part of the anterior root of the 
main ganglion. Both parts entered the truncus maxillaris. 
Whether they represent the two bundles into which their root 
separates on entering the brain, or only one of those bundles, 
was not determined. The fibres arising from the postero- 
dorsal portion of the main root disappeared gradually as they 
approached the truncus maxillaris. The transverse bundle in 
the ganglion is called by Strong in Rana (No. 121, p. 135) the 
motor bundle of the portio minor of the trigeminus. 
The orbital vein, which lies at first anterior to, and internal 
to, the upper, anterior part of the ganglion, runs outward, 
downward, and backward under that portion and its root, and 
issues on the dorsal surface of the median and posterior portions 
of the ganglion. It thus pierces the main ganglion under its 
