624 ALLIS. [Vou. XII. 
laris. The fibres of the anterior of the two roots ran forward 
and outward, and supplied first the macula utriculi, which lay 
immediately above the ganglion of the root, and then the crista 
ampullae anterioris. The fibres of the posterior of the two 
roots ran outward and slightly forward and supplied the crista 
ampullae externae. 
The main posterior root, or ramus cochlearis, and its gang- 
lion were always much more distinctly separated from the other 
two roots and their ganglia than those two roots and ganglia 
were from each other. 
As the ramus ampullae posterioris passes outward and back- 
ward between the lagena and the sinus posterior it passes 
immediately above the root of the nervus glossopharyngeus, 
which runs at this point almost directly outward. The two 
nerves here come into intimate relations and there is always 
an interchange of fibres, the exact nature of which I have been 
unable to satisfactorily determine. To the best of my observa- 
tion some of the fibres of the acusticus join the dorsal or sense- 
organ root of the glossopharyngeus, but whether they run 
distally or proximally in that root I am unable to say. Distal 
to this point the dorsal root of the glossopharyngeus is sepa- 
rate and distinct from the remaining much larger portion of 
the root of that nerve, lying immediately behind it and having 
an entirely separate ganglion, as already described in my earlier 
work (No. 3, p. 516). While passing inward under the acusti- 
cus, through the narrow space between the lagena and utriculus, 
the two roots of the glossopharyngeus are much flattened, and 
the large fibres of the dorsal root were always lost in my sec- 
tions. Median to the acusticus, after passing under it, they, 
however, reappear in the sections and receive an important bun- 
dle of fibres from the acusticus, the bundle being apparently as 
large as the dorsal root of the glossopharyngeus itself. Proxi- 
mal to this anastomosis the dorsal root still persists, lying in its 
accustomed position immediately behind and in contact with 
the main root of the glossopharyngeus. This latter root, in a 
14 mm. and 20 mm. specimen of which I had excellent sections, 
then passed through the root of the nervus lineae lateralis, 
near its ventral edge, and there was again an interchange of 
