642 ALLIS. FVOL. Xt: 
backward, become connected with a second, pretrematic, series 
of ectodermal thickenings. From both these thickenings 
ganglia arise, the first or epibranchial ganglion being common 
to the post- and pretrematic branches of the nerve, while the 
second or pretrematic ganglion is connected with the pretre- 
matic nerve alone. In Petromyzon no ectodermal sense organs 
of any kind arise in connection with the epibranchial ganglia 
(No. 67, p. 30). If this be true for other fishes, terminal buds 
must arise in connection with the pretrematic ganglia and 
ectodermal thickenings, if their origin and development is sim- 
ilar to that generally accepted for the organs of the lateral 
line. They would, therefore, seem to be the rudimentary 
sense organs found by Froriep on the facialis, glossopharyn- 
geus, and vagus in calf embryos, and as the eye belongs, 
according to Kupffer (No. 67, p. 50), to the line of epibranchial 
ganglia, the lens and its nerve, the ciliaris brevis, may belong 
to the pretrematic series. Sedgwick says (No. 114, p. 97) 
that the profundus ganglion, from which the ciliaris brevis and 
other ciliary nerves arise, is, when it is first laid down, in 
contact with the ectoderm. 
In short, the nerve fibres arising from the fasciculus com- 
munis tract seem destined in large part, if not in whole, to the 
supply of terminal buds, as Strong (No. 121) has suggested 
might be the case. The fibres so arising may issue from the 
brain as a separate and distinct root, on which a separate and 
distinct ganglion is found; they may issue in part as com- 
ponents of certain nerves with the roots of those nerves, and 
in part as a separate root which becomes immediately more or 
less fused, it and its ganglion, with other roots and ganglia ; 
or they may apparently issue entirely as components of certain 
nerves. 
To the first category belongs, apparently, Protopterus ; to 
the second, Amia, Rana, and many other fishes and amphibia ; 
to the third, birds, judging from Brandis’ descriptions (No. 12, 
p. 539, and-No. 13, p..647), for he finds the fibres of the 
funiculus solitarius issuing with the facialis and glossopharyn- 
geus, and possibly also with the vagus, and the funiculus soli- 
tarius of higher vertebrates corresponds, according to Strong 
