812 FOOT. [VoL. XII. 
matic network of the pronucleus, while their nucleoli grow and 
fuse with one another. Fig. 5 shows several small vesicles 
and a few relatively large vesicles, the result of the fusing of 
two or more small ones. Just before the vesicles appear, the 
chromosomes select the methyl green stain, but as soon as 
the vesicles are formed we have a tiny, green nucleolus sur- 
rounded by ye//ow chromatin, this clearly suggesting that the 
nucleolus is a substance which a moment before was contained 
in the chromosomes. The chromatin which at first surrounds 
each vesicle continues to select the orange stain until it has 
again assumed the form of chromosomes. Fig. 6 shows the 
male pronucleus forming in exactly the same manner as the 
female pronucleus, the head of the spermatozoon breaking up 
into vesicles similar to those of the young female pronucleus. 
Vesicles entirely similar to those found at the telophase of 
the second maturation spindle are found also at the telophase 
of the first maturation spindle, though I have never seen any 
evidence of their development into a resting nucleus. 
Sperm Granules, Fig. 2.— The sperm granules are not con- 
stant structures. In eggs found during the height of the 
breeding season (when they are less likely to present ab- 
normalities), the sperm granules are either not present at all 
or are relatively insignificant in both size and number, and 
in such cases they are often found near the posterior end 
of the head of the spermatozoon. Near the close of the 
breeding season, however, when few normal eggs are found 
(sometimes only one in a cocoon that contains fitty eggs, show- 
ing various degrees of degeneration), 

at this season nearly 
all the eggs having one or more fertilization cones contain 
relatively many and large sperm granules. These granules 
appear to be formed at the expense of some of the surrounding 
substance, and such preparations as are represented by Fig. 2 
suggest that the substance sacrificed to the formation of these 
bodies is archoplasm, for there is relatively very little archo- 
plasm at the side of the cone occupied by the sperm granules. 
The suggestion that these granules are metamorphosed 
archoplasm raises several questions that I am entirely incom- 
petent to answer. Wherein do they differ from apparently 

