1870. ] ANATOMY OF THE PRONGBUCK, 337 
animal’s physiological relations or affinities, which, if not weighty, at 
all events have their value. 
There being neither a suborbital gland (crumen), osseous fossa for 
the same, nor inguinal sacs and pores, points consequentially to forms 
exhibiting a kindred build. The total deprivation of the former 
shows but a remote alliance to the cervine structural peculiarities— 
though it does not necessarily constitute it an Antelope ; for a large 
section of the Antelopes possess a crumen. But a minor series, 
chiefly of the Goat-like forms, have it not ; and to this group, then, 
the Prongbuck would be linked—a union which is strengthened by 
the fact that such Antelopes as are distinguished by the absence of 
inguinal sacs and pores come under the same group. 
Among Ruminants the Chamois is noted as having a glandular 
sac which opens behind the ear, though some authors indicate Pro- 
capra as having a postcorneal sinus. In the possession of a sub- 
auricular skin-gland Antilocapra announces organic relation. More- 
over the rank hircine odour from the above, as well as the circum- 
stance of a glandular tail-patch, decidedly point to Capra. 
In brief, were the place of the Prongbuck to be assigned by the 
number, situation, and secretion of its skin-glands alone, I should 
without hesitation rank it among or close to the Goats. 
Casting a glance among the viscera and other internal soft struc+ 
tures of Antilocapra, the subjoined points demand attention. There 
being a gall-bladder severs it from the Cervidz and allies it with the 
Bovide. The stomach having four fully developed cavities, there 
being no water-chambers in the rumen, and no ileo-czeal gland ex- 
clude it severally from the Tragulide, Camelidee, and Giraftide, 
The tongue might either belong to a Goat, an Antelope, or a Stag, 
though probably more like that of the two former than the latter. In 
the non-development of Cowper’s glands, in the manner of the termina- 
tion of the vasa deferentia, and in the bluntish form of the glans penis, 
the generative organs denote consanguinity with the Deer, where 
such structural conditions preponderate in the group. The con- 
struction of the larynx has a sort of medium tendency of divergence, 
the general type being cervine, though the short upper cornua would 
rather signify kindred with the Chamois and some Antelopes. As 
to the liver, its having a gall-bladder takes the Prongbuck away 
from the Cervi, which have no such reservoir. 
Regarding dental characters, I can see no obvious distinction be- 
tween the teeth-pattern of the Cabrit and the family Antilopide. 
The appendicular skeletal segments show long limbs of a strong 
but manifestly fine and delicate construction, such alone as belong to 
the light fleeting Antelope tribe ; for in all Deer, Goats, and Sheep 
of similar size, relatively much stouter leg-bones pervade. 
In groups of the Ruminants there is considerable variation in the 
sternum, according as its component pieces are broad or narrow and 
the preesternum compressed or flat. As a rule the elementary parts 
are broader in Deer than in Antelopes ; and in this moderate breadth 
to length the Prongbuck follows the latter. It also agrees with 
them im the form of its pelvis. 
