506 DR. J. MURIE ON HYZNA BRUNNEA. 
walls, of some thickness, are strictly glandular. The matrix in which the glands 
are imbedded is a cellulo-fibrous material, through which the glands pierce by pores 
opening into the interior of the sac itself. The three anal saccular glands on each side 
of the rectum, though to some extent separate from each other, have nevertheless close 
union, lying adpressed the one to the other, so as to represent but a single body some- 
what larger than a walnut, and, like that fruit, rough and warty outside (wide Pl. LXIII. 
fig. 2, dotted lines). The above description applies to the dissected condition ; for exte- 
riorly the anal glandular chambers are enclosed in a thick, dense, muscular mass, the 
compressor glandule (c.g); and this increases the bulk of the bodies to the size of a 
small orange, as shown in fig. 1 of the Plate. 
The glandular secretion is a darkish unctuous substance, with a peculiar rancid 
butyraceous odour, as Daubenton faithfully notes. It collects in quantity in the afore- 
said cavities and passages, and doubtless exudes under pressure of the muscular appa- 
ratus, as, I believe, during sexual excitement. I have myself observed the Hyznas in 
the Gardens during such condition rub the boards of their den with the hinder end of 
their body, leaving behind traces of the odoriferous anal-gland secretion. The Hyena's 
scent-bags agree in several respects with such developments in some of the Viverridee, 
and thus point to relationship with that family’. 
Professor Kaup, a long time ago’, attempted to divide the Hyznas generically accord- 
ing to their dentition, and laid stress on some and not others possessing an anal sacculus. 
The spotted species he separated as a genus Crocuta. Mr. Busk*, in a recent paper of 
his, alludes (p. 60) to the above division, and to Dr. Falconer’s proposal* to adopt the 
term Euhyena for the other section. These writers have each apparently taken it for 
granted that H. maculata is deficient in anal glands. 
Mr. Busk says, in a footnote (p. 71), “* Having lately examined a living H. crocuta in 
the Zoological Gardens, I can confirm Kaup’s statement that no trace of a pouch 
between the root of the tail and the anus exists, at any rate in the male of that 
species.” 
I myself was present and aided in the above most hazardous undertaking ; and cer- 
tainly it did seem, on hasty and tactile search, that a postanal pouch was absent. Since 
then I have had reason to change my opinion, inasmuch as in a very young dead male 
spotted Hyzena which passed through my hands, sure enough there was an ample post- 
anal pouch and glandular development connected therewith (see fig. 2, p. 507), just as 
in the female H. drwnnea. 
On what grounds Professor Kaup’s assertion is made I know not, and that a postanal 
1 Since this paper was read, Professor Flower has given us an account of the anatomy of Proteles cristatus 
(P. Z. 8. 1869, p. 494); and therein mentions its having a supraanal follicle and lateral glandular sacs. 
? Isis, 1828, p. 1144. 
4 « Remarks on the cranial and dental characters of the existing species of Hyena,” Journ. Proc. Linn, Soc. 
vol. ix. 4 Paleontological Memoirs, vol. ii. p. 464. 
