20 The thallosc Ilepaticae of the Juan Fernandez Islands. 



By 



ALEXANDER W. EVANS. 



With six text figures. 



The thallose Ilepaticae include the Marchantiales, the Jungermanniales 

 Anacrogynae, and the Anthocerotales. The Marchantiales are represented in 

 the Juan Fernandez islands by the Rebouliaceae and the Marchantiaceae; the 

 Jungermanniales Anacrogynae by the Riccardiaceae, the Monocleaceae, and the 

 Pelliaceae; and the Anthocerotales bj' the Anthocerotaceae, the only family of 

 the order. 



The earliest collection of Hepaticae from the Juan Fernandez islands was 

 made in 1830 by the Italian botanist, C. G. Beutero. It includes several 

 thallose species, the most conspicuous of which is Marchantia Berteroana Lehm. & 

 Lindenb., which was described as new in 1834.^ This robust species, one of 

 the largest members of the genus, is now known to be widely distributed in 

 the higher latitudes of the Southern Hemisphere. 



In the following year MoNTAGNE published a full account of BerteRo's 

 cellular cryptogams under the title: »Prodromus F'lorae Fernandesianae. Pars 

 prima, sistens enumerationem plantarum cellularium quas in Insula Juan Fer- 

 nandez a CI. Bertero collectas describi edique curavit C. Montagne, D. M.»^ 

 In this paper he reported three additional thallose hepatics from the islands, 

 namely: Jiingermannia furcata L., J. pinnatifida Nees, and J. rhizoloba Schwaegr. 

 The specimens upon which the first two records are based have not been 

 available for study. It is hardly probable, however, that the plants referred to 

 J. furcata represent Metzgeria furcata (L.) Dumort. of northern regions. It 

 is much more likely that they would now be included under M. decipieiis 

 (Massal.) Schififn. & Gottsche, and the plants referred to J. p27inatifida perhaps 

 represent Riccardia brevvanwsa (Steph.) Evans. The plants referred to J. 

 rhizoloba were made the type of Syniphyogyna HocJistetteri Nees & Mont, in 

 1836.^ They proved to be distinct from the true J. rJiizoloba of Bourbon, now 

 known as Symphyogyiia rhizoloba (Schwaegr.) Nees. In 1850 MoNTAGNE re- 

 ported the cosmopolitan Marchantia polymorpJia L. from Juan Fernandez, on 

 the basis of one of Bertero's specimens.^ 



1 In Leh.m.axx, Pug. PI. 6:21, 49 (1854). 



2 Ann. Sci. Nat. II, 3:347—3)6; 4:86—99 (183)). 

 ' Ibid. II. 5:68 (1836). 



* In Gay, Hist. Fis. Polit. Chile 7: 508 (1850). 



36—30579. The Xat. Hist, o/ Juan Fernandez and Easter Isl. Vol. II 



