C. SKOTTSBERG 



The distribution of Lagenophora clearly testifies to its Antarctic origin: 3 

 Magellanian species, one of them also on Tristan da Cunha and one on Masafuera, 

 7 in New Zealand, 2 in Australia, one of these north to the Philippines, i in Fiji 

 and 3 in Hawaii, the four Pacific ones connected with L. Billarduri CdiSS. (Austral.), 

 whereas the Magellanian species point toward L. puuiila Cheescm. d^ud pe/iolata 

 Hook. fil. (New Zealand). 



In Erigeron we meet again with a large boreal genus strongly represented 

 along the Andes and ranging south to Patagonia, Fuegia and Falkland; 135 species 

 are reported from Eurasia, inch India, 345 from North America, 35 from Centr. 

 America and the W. Indies, 95 from S. America, 10 from Africa, 8 from Australia 

 and a single species from Oceania (Rapa), E. rapensis F. B. H. Brown, and this is 

 compared with our island species. To judge from the description it has the same 

 habit as these, a shrub about 3 dm tall with the leaves in terminal rosettes. In 

 Hawaii the genus is represented by Tetraviolopiiini, which is very close to Erigeron; 

 ViERHAPPER {22g. 1 82) suggested that the Juan Fernandez species came near the 

 Hawaiian E. lepidotus Less., which is now referred to Tetraniolopiuni, and that we 

 ought to look for relatives among the Andine and Mexican forms. Among our 

 island species E. rupicola differs much in habit from the rest, but all belong together 

 and most likely represent a special branch of the Andine Erigeroji flora which, in 

 its turn, comes from a boreal stock. 



To what I said above about Guaplialiinn spidforuic nothing can be added 

 at present. Chile is well provided with poorly limited species related to G.pur- 

 pureuni L.; most of them are badly known. Their boreal parentage can hardly 

 be doubted. 



Abrotanella resembles LagenopJiora in its distribution, but is absent from Oce- 

 ania; the majority inhabits New Zealand with its subantarctic islands (9 sp.), i is 

 found in Victoria, 2 in Tasmania, i in New Guinea and 5 in S. America (W. 

 Patagonia to Fuegia and Falkland); in addition, one is found on Rodriguez I. 

 The genus is, as it were, tricentric and its Antarcto-tertiary character indisputable. 



The concentration of isolated arborescent Compositae in the Pacific was em- 

 phasized by Bentham; the main groups of the family are represented among them, 

 and the accumulation of endemic genera in Hawaii and Juan Fernandez has led 

 to much discussion. The enormous development and differentiation in the family 

 on American soil is an undeniable fact, but simply to derive the endemic Pacific 

 genera from America as did GUPPY {121) does not seem possible. Speculations (comp. 

 f. inst. Setchell 2ig) led back to Antarctica, but not until Brachionoslyluui was 

 discovered in New Guinea and found to be nearly related to Robvisonia of Juan 

 Fernandez were we able to stand on tolerably firm ground. 



With regard to Ceiitaiirodendron, to which Yunquea seems to be related, the 

 situation is different. Centaurea and all the genera of Centaureinae belong to the 

 Old World with the exception of a single species in N. America and a few in 

 the Andes, south to Centr. Chile. The group ranges over Europe with a strong- 

 hold in the Mediterranean, the Orient, Ethiopia and through Centr. Asia to Japan. 

 The Chilean species of Centaurea belong to the Plectocephali, but Cenfauroden- 

 dron differs not only from this section but from all in sex distribution [243). It 



