Tribe 12. ANTHEMIDEAE 
PER AXEL RYDBERG 
Heads radiate or discoid. Ray-flowers when present pistillate or rarely 
(among ours only in Maruta) neutral; corolla ligulate, or tubular and 2-4- 
lobed, or (in Lancisia, Gymnostyles, and Soliva) wanting. Disk-flowers 
hermaphrodite and fertile, or in some species of Artemisia sterile, with actino- 
morphic 4- or 5-lobed corollas. Involucral bracts herbaceous with dry 
scarious margins and tips. Receptacle with or without chaff, from nearly 
plane to high-conic. Anthers inserted at the rounded or obtuse base, without 
basal appendages, the connective produced into a short obtuse or a longer 
subulate tip. Style-branches of the hermaphrodite flowers short, truncate 
and hispidulous, fimbriate at the apex, without appendages, or in some species 
of Artemisia the style undivided; style-branches of the pistillate flowers short, 
oblong, obtuse or truncate at the apex, rarely filiform. Achenes small, cylindric- 
oblong or more or less flattened, often unequally ribbed. Pappus wanting or a 
crown of united squamellae, seldom stronger and unilaterally developed. 
Leaves usually alternate and often dissected. 
Receptacle with chaff, this usually consisting of membranous, more or less 
concave elongate scale-like paleae. 
Style-branches with short appendages; achenes large, obpyramidal 
(See Leucampyx in Tribe HELENIEAE, page 55). 
Style-branches without appendages, in the hermaphrodite flowers trun- 
cate and hispidulous, fimbriate at the apex; achenes small, oblong 
or cylindro-prismatic. : 
Heads usually radiate; disk-corollas with a cylindric tube and elon- 
gate-funnelform throat; anthers with short ovate tips. 
Achenes more or less obcompressed, sub-rhombic or obliquely 
triquetrous in cross-section; tube of the disk-corollas more or 
less flattened. 
Heads small, corymbose-paniculate; involucre campanulate 
to hemispheric; ligules few, 5-12, short and broad; pappus 
wanting. 1. ACHILLEA. 
Heads large, solitary at the ends of the branches; involucre 
saucer-shaped; ligules 10-30, elongate, linear; pappus of a 
short membranous crown. 2. Cora. 
Achenes terete, cylindric or ellipsoid; tube of the disk-corollas 
cylindric. 
Ray-flowers fertile; paleae of the receptacle membranous, 
subtending all disk-flowers. 
Tube of the disk-corollas produced into a spur-like append- 
age at the base; ligules in ours white with yellow bases. 3. ORMENIS. 
Tube of the disk-corollas not appendaged at the base; 
ligules white or wanting. 4. ANTHEMIS. 
Ray-flowers sterile; paleae of the receptacle subulate, stiff, 
subtending only the upper disk-flowers. 5. Maruta. 
Heads discoid, numerous; corollas funnelform without a distinct tube; 
anthers with subulate tips. 18. ARTEMISIASTRUM. 
Receptacle without chaff. 
Style-branches with conic or suhulate appendages (See HyMENOPAP- 
PANAE in Tribe HELENIEAE, page 43). 
Style-branches without appendages. 
Style-branches linear-filiform (See PALArOxIANAE in Tribe HELEN- 
IEAE, page 58). 
Style-branches at least of the disk-flowers short and truncate at the 
apex. 
Heads radiate, with well-developed usually flattened ligules; 
anther-tips ovate, mostly obtuse. 
VoiumeE 34, Part 3, 1916) 217 
