Part 3, 1916] CARDUACEAE: ANTHEMIDEAE 227 
wide, closely and finely serrate; heads few in open simple or compound corymbs; involucre 
hemispheric, villous, 4-5 mm. high, 8 mm. broad; bracts 25-30, in about 3 series, the outer 
lanceolate, the inner oblong, all acutish and with pale margins; ray-flowers 8-10; ligules 
white, 4-5 mm. long, suborbicular, round-lobed; disk-flowers 50-75; corollas 3 mm. long; 
achenes 2 mm. long, broadly wing-margined. 
TYPE LocALiTty: Temperate Europe. 
DIsTRIBUTION: Labrador to New York, Missouri, and Michigan; adventive or naturalized 
from northern Europe and Asia. 
ILLUSTRATIONS: Curt. Fl. Lond. 5: pl. 60 [2: pl. 176]; Engl. Bot. pl. 757; Fl. Dan. pl. 643; 
Sv. Bot. fl. 207; Hayne, Arzn. Gew. 9: pl. 44; Baxter, Brit. Bot. pl. 36; Reichenb. Ic. Fl. Germ 
a: pl. SBE Coste, Fl. Fran. f. 1969; Fl. Deuts. ed. 5. pl. 3016; Britt. & Brown, Ill. Fl. f. 3982 
Bee fe 4552- 
’ 
2. COTA J. Gay; Guss. FI. Sic. Syn. 2: 866. 1844. 
Annual or perennial herbs. Leaves alternate, pinnatifid or bipinnatifid. Heads solitary 
at the ends of the branches, usually radiate. Involucre saucer-shaped; bracts in about 3 series, 
rather firm, the outer somewhat shorter. Receptacle convex, chaffy; paleae membranous, 
cuspidate. Ray-flowers 10-30, pistillate and fertile; corolla-tube obcompressed and acutely 
margined, at least below; ligules white or yellow. Disk-flowers hermaphrodite and fertile; 
corolla-tube obcompressed, at least equaling the elongate-funnelform throat; lobes 5, lanceo- 
late, spreading. Anthers with ovate acute tips. Style-branches in both kinds of flowers 
truncate at the apex. Achenes obcompressed, somewhat triquetrous, the lateral angles 
prominent, the two faces each about 10-nerved. Pappus a short membranous crown, in ours 
entire. 
Type species, Anthemis Cota L. 
Ligules white; leaves bipinnatifid. 1. C. altissima. 
Ligules yellow; leaves pinnatifid. 2. C. tinctoria. 
1. Cota altissima (L.) J. Gay; Guss. Fl. Sic. Syn. 2: 867. 1844. 
Anthemis altissima L. Sp. Pl. 893. 1753. 
A tall leafy annual; stem 3-8 dm. high, terete, glabrous or nearly so, branched; leaves 
3-6 cm. long, obovate in outline, bipinnatifid into linear, entire or toothed, spinulose-tipped 
divisions, sparingly hirsutulous; heads solitary at the ends of the branches, radiate; involucre 
8 mm. high, 15-18 mm. broad; bracts pilose, the outer somewhat shorter, ovate, acute, the 
inner oblong, obtuse; receptacle convex; paleae obovate-spattilate or cuneate, abruptly cuspi- 
date, equaling the disk-flowers; ray-flowers 15-20; ligules white, about 12 mm. long, 3-3.5 
mm. wide; tube somewhat obcompressed below, longer than the throat; achenes 3 mm. long. 
TyPE Loca.ity: Italy. 
DIsTRIBUTION: Oregon; adventive from the Mediterranean region. 
ILLusTRATION: Coste, Fl. Fr. f. 1959. 
2. Cota tinctoria (I,.) J. Gay; Guss. Fl. Sic. Syn. 2: 867. 1844. 
Anthemis tinctoria L. Sp. Pl. 896. 1753. 
Chamaemelum tinctorium Schreb. Spicil. Fl. Lips. 145. 1771. 
A leafy perennial; stem 3-6 dm. high, more or less hirsute-villous, somewhat branched, 
striate, often purple-tinged; leaves 2-5 cm. long, pinnatifid, with a winged and toothed rachis, 
hirsute-villous; divisions oblong, and cleft, incised, or sharply serrate, with cuspidate teeth; 
involucre saucer-shaped, 5-6 mm. high, 12-18 mm. broad; bracts villous, lanceolate, acute, 
the outer slightly shorter; disk convex; paleae linear-lanceolate, aristate-tipped, equaling the 
disk-flowers; ray-flowers 20-30; ligules yellow, 12-15 mm. long, 3-4 mm. wide; disk-corollas 
3.5—-4 mm. long; tube fully equaling the throat; achenes 2 mm. long. 
Tyre LOCALITY: Sweden. 
DisrrisuTion: Maine and Quebec to New Jersey and Ohio; naturalized from Europe. 
IntustRations: Fl. Dan. pl. 741; Sv. Bot. pl. 62; Engl. Bot. pl. 1472; A. Dietr. Fl. Bere ee 
641; Reichenb. Ic. Fi. Germ. 16: pl. 1010, f.1; Fl. Deuts. ed. 5. pl. 3031; Coste, Fl. Fr. f. 
Britt. & Brown, Ill. Fl. f. 3987; ed. 2. f. 4559. 
