1889.] FAT-BODIES OF THE SAUROPSIDA. 603 
Page 
V. On certain Subcutaneous Fat-deposits .............csceseeeeeeeer eee 609 
VI. On the fatty “Spleen” of the Crocodiles................sssssseereeeee 610 
VII OoniGhisiongs vic scccetnse teas veces tedocedsscaessasbarsbcoeedevesniwaskoeseees 611 
VT. Explanationtof tho Plates.cs. <<. ..s.c.0~sscvaeh~ stones cadestasadtuettoes 612 
I. InrropucrTory. 
The conditions under which the investigations of which this paper 
gives the results were commenced and carried on were stated in the 
introduction to my paper upon the Subdivision of the Body-cavity, 
read at the last meeting (see above, p. 452). 
II. On THE Revations oF THE FaT-BoDIEs OF THE SAUROPSIDA, 
AND ON CERTAIN POINTs IN THE ANATOMY OF MoniToRs. 
The fat-bodies referred to are those which, as is wel! known, 
occur in Lizards on the course of the “ pelvic’’ veins, and of more 
or less of the anterior-abdominal vein. ‘The vessels named, with 
their tributaries, take away the blood, which is brought to the fat- 
bodies by large branches from the anterior of the two pairs of 
arteries that supply the hind limbs, and which I take to be homo- 
logous with the femoral arteries of birds. 
Corresponding fat-bodies are very conspicuous in the Snakes (ef. 
figs. 8, 9, & 10 ¢.a), where, as in the Snake-like Amphisbeenide, 
they extend from the cloaca to the hinder margin of the liver (ef. 
figs. 4, 5,6, 7). The figures of sections of Adder and embryo Grass- 
Snake show that, when the fat is well developed, the peritoneal 
cavity of Snakes may be much restricted by reason of the fact that 
the kidneys and fat-bodies lie outside it. The latter occur in the 
Crocodiles, but, as described below, the fat-bodies here referred to 
are in these animals more lateral in position than in Lizards; and in 
the case of the birds, the fat-laden “omentum,” or transversely 
expanded ventral ligament of the stomach, is, I think, obviously 
comparable, so far as its fat is concerned, to the similar fat-laden 
ventral ligament in such forms as the Amphisbeenide and Snakes, 
where the fat extends forwards as far as the liver. 
The Chelonia are the only order of the Sauropsida in which I 
have not observed these structures well developed, but there appear 
to be traces of them in Emys europea’. 
In many Lizards (ef. fig. 11) these fat-bodies, pushing the 
peritoneum before them, bulge into the body-cavity ; and, lying on 
the course of the large vessels ventral to the (once respiratory 
allantoic) bladder (ef. figs. 7 & 12) and the alimentary canal, into 
the ventral ligament of which they in some forms (Amphisbxnidz, 
fig. 4) obviously extend, they may form paired masses quite as con- 
spicuous in the posterior part of the abdominal cavity as are the 
liver-lobes in the anterior half*. 
1 T have only examined in this connection some half-dozen specimens of 
Emys and Testudo, and those not large ones. 
* These lie, of course, ventral to the alimentary canal and lungs. Passing 
oyer the important difference that no branching system of tubules extends from 
40* 
