236 PHILOSOPHICAL NOTES ON 



the teeth of leaves as remnants of aborted sori, the originals of 

 which might be such as we see on the tips of the teeth of Adian- 

 turn and others. Indeed, the teeth of leaves receive a separate 

 vein or fibro-vascular bundle in the same way that the Adiantum 

 sorus receives the termination of a vein. 



The two anther cells, to my mind, cannot but be homologous 

 with the two petiole glands or of others like them, the leaf -blade 

 being suppressed. When the stamen reverts to a leaf, as in Fig. 83, 

 the anther or anthers return to their position of glands, although, 

 of course, they may remain poUiniferous, owing to incomplete 

 reversion. 



Curiously enough, after conceiving all this, I accidentally came 

 across an adumbration of what I have been trying to work out. I 

 have likened the anthers to the glands on the petiole of the cherry- 

 leaf, and this is what Asa Gray (" Structural Bot.") states in a note 

 to page 254 : " Sachs takes the filament with the connective to be the 

 homologue of the whole leaf, and the anther cells as appendages* ; 

 others in likening the anthers to glands, adopt a similar view " ! 



According to my view, we would have the amalgamated glands 

 of the cherry petiole teeth-glands, pollen grains, and ovules, all 

 homoloo-ues. Then I have shown that teeth are no other than 

 abortive branches, so that the marginal buds of the Bryophyllum. 

 take their place as branchlets of the teeth, and become homologous 

 with axillary buds. We have thus a whole string of homologies — 

 axillary and marginal buds, glands of the cherry petiole, of the 

 peach and other leaves, marginal tooth-glands of leaves, pollen 

 grains, and ovules, with their plumule or seed-bud. All these 

 become homologous according to this view. This is exactly what 

 I hinted at on pages 211 and 246 of the " Cultivated Oranges and 

 Lemons of India and Ceylon." 



I do not enter into the microscopical structure of all these 

 ijarts because I know that in the cell there is a nucleus and 

 crranules within the nucleus another nucleus and granules, and so 

 on. Not only is the anther cell, in its incipience, potential pollen 

 grains but the ovule cell is a potential whole tree, with its stem, 

 roots leaves, stamens, pistils, and again ovules. To each little step 

 in this evolution, histologists appear to have given a different name. 



* What can appendages mean morphologically ? 



