286 PHILOSOPHICAL NOTES ON 



seaweeds, however, they are ramelli, or small .branches. When 

 atrophy occurs, once important organs may not only become 

 epidermal, but often quite extinct. 



I know that pollen grains are generated by fours in the 

 interior of the anther nipple in mother cells, nevertheless, they 

 eventually become superjicial, and are shed exactly like the 

 globular hairs on the leaves of Chenopodiimi petiolare and other 

 Chenopodiums. 



The reader should not forget that I start with the 

 notion that every cell is a potential reproductive organ. 

 He should also take note of the fact that Gr. O. Bower found 

 glandular hairs in the intercellular spaces of Aspidium* So that 

 glandular hairs are not only produced on the exterior, as an out- 

 come of the epidermis, but also in the interior of the parenchyma ! 

 This is not surprising, when we think that there is no good reason 

 why external cells should be morphologically distinct from internal 

 cells. The fact of their being outside of the internal ones, and 

 thus forming a sort of covering to the inner ones, and the fact of 

 their having been called epidermis, do not alter their nature 

 morphologically. This remains identical with that of the ancestral 

 monophyta, from which they descended, although their function 

 may be changed. 



Le Maout and Decaisne, " System of Botany," p. 139, say that 

 " True glands difPer from glandular hairs only in projecting 

 slightly, or not at all, above the epidermis ; they pass insensibly 

 into each other, as in glandular roses." 



In the Droseras we have these glandular hairs developed into 

 large sensitive tentacles. And it is in these plants that we find 

 light thrown on the real nature of glandular hairs, which are found 

 in many plants, although in the moss-roses the different parts are 

 crowded with them. 



Prof. Henslow (" Structure of Flowers," p. 307) says :— " The 

 tentacles of Drosera, as is well known, are not epidermal trichomes, 

 but structures issuing from branches, arising from the libro- 

 vascular cords of the leaf, and therefore strictly homologous with 

 the * funnels * on cabbage leaves."f In a note he adds : — " The 



* " Practical Botany," p. 293. 



f May not these " funnels " be more appropriately homologized with 

 the leaflets on the midrib of Delesseria coriifolia. Fig. 12. 



