PISTILLODY. 307 
shows one of the intermediate organs from these 
flowers, in which half the structure seems devoted to 
the formation of ovules, while the other half bears a 
one-celled anther. Lindley’ has also 
described a case of this kind in a 
species of Amaryllis. 
In Savifraga crassifolia it sometimes 
happens that mixed with the stamens, 
and originating with them, are a 
number of distinct and _ perfectly 
formed carpels, wholly separated from 
the normal carpels, m the centre of 
the flower. In this particular in- 
stance there is usually no interme- 
ciate condition between the stamen 
and the pistil. Guillemin’ also de- 
seribes a transformation of the stamens Fre. 164.—Struc- 
into carpels in Huphorbia esula. ture half anther, 
When the anther is involved it may *#!f ca"pel, ilvan. 
be only partially so, or almost the whole organ may be 
transformed. As instances of very partial change 
may be cited the passage of the connective into a 
stigma in Thalictrum minus, or the passage of the 
points of the anthers into imperfect styles in some 
species of bamboo.’ 
In Rosa arvensis similar transformations have been 
observed of a slightly more complex character than 
those just mentioned, and passing into more impor- 
tant changes, especially to the formation of pollen 
within ovules, formed on the edges of an open car- 
pellodic anther (see p. 186). 
Mr. Berkeley has recorded an analogous case in a 
gourd in which the stamens bore numerous ovules (p. 
200), and. Baillon describes another gourd in which cer- 
tain fleshy appendages surrounding the androecium 
were provided with ovules.* 
1 «Theory of Horticulture,’ ed. 2, p. 82. 
2 «Mém. Soc. Hist. Nat. Paris,’ i, 16. 
* Gen. Munro, ‘ Trans. Linn. Soc.,’ xxvii, p. 7. 
* «Bull. Soe. Bot, Fr.,’ 1857, p. 21. 

