Fink: LYCOPERSICUM ESCULENTUM. 641 



cut off the stigma entirely just as the bud was opening. It ap- 

 pears from this that the pollen will occasionally germinate on 

 the cut end of the style after the stigmatic surface is removed, 

 hence my failure to get one-sided fruit in some instances when 

 one side of the stigma had been removed. I also tried pollin- 

 ating on one side only and got one-sided fruit as a result. 



The Handsomest and Best tomato was experimented with as 

 follows: Twenty-five flowers were castrated as fast as they 

 began to open and were pollinated with pollen from as many 

 different varieties of tomatoes. Fifteen mature tomatoes re- 

 sulted and four more aborted after beginning to grow, proba- 

 bly from insufficient pollination. The other six severed their 

 connection with the vine at the joint of the pedicel. The 

 number of failures was no greater than in flower clusters that 

 were marked and left to natural processes of pollination. In 

 the experiments each stigma was pollinated only once. Later 

 I tried crossing the Tree tomato, pollinating each castrated 

 flower three days in succession. The result was fifteen toma- 

 toes from sixteen castrations. 



For the purpose of studying the offspring of a cross the Tree 

 tomato was crossed with the Plum-shaped Yellow, the Pear- 

 shaped, the Red Cherry and Yellow Cherry varieties, as these 

 represent the principal differences between the Tree tomato 

 and other varieties as to shape and size of vine, fruit and leaves. 

 The seeds were planted the next year, and I could not see that 

 the vines, leaves or fruit showed a tendency to imitate the 

 plants bearing them more than that from which the pollen was 

 taken for pollination. The fruit was about intermediate be- 

 tween the two plants crossed. However, more observation 

 than I was able to make is necessary to settle this point. 



It has been supposed that crossing produces a difference in 

 the size, shape and color of the tomato resulting from the cross. 

 The largest tomato produced by crossing the Handsomest and 

 Best was from a cross with the yellow Cherry tomato, which was 

 my smallest variety. The crosses with the Large Yellow, 

 Yellow Cherry and Yellow Plum-shaped varieties produced 

 fruit as deep red as those not crossed. A cross with the Yel- 

 low Plum-shaped tomato produced a tomato which happened 

 to be more flattened than the Handsomest and Best tomatoes 

 usually are instead of a longer one. Close observation of a 

 large number of tomatoes from crosses seems to show that the 

 size of the fruit is slightly increased and that the crossed 



