Family 10. MALPIGHIACEAE 



By John Kunkel Small 1 



Shrubs, trees, or partially woody herbs, or vines with reclining, trailing, or 

 twining stems. Leaves usually opposite, with stipules at the base of the 

 petiole or on it ; blades narrow or broad, entire, toothed, or lobed, often with 

 glands on the margins or on the back. Flowers usually perfect, solitary or 

 variously borne in terminal or lateral clusters. Cleistogamous flowers with 

 more or less rudimentary organs accompany the other flowers in some genera. 

 Calyx of 5 sepals, some or all of which bear sessile or stalked glands, or occa- 

 sionally all are glandless. Corolla white or usually variously colored, of 5 

 unequal petals which are usually manifestly clawed, the blade undulate, toothed, 

 fimbriate, or lobed, and often concave. Androecium of 5 or 10 anther-bearing 

 stamens, or sometimes fewer ; filaments united at the base or higher up, rarely 

 distinct ; anthers narrow or broad, the connective often large. Gynoecium of 

 typically 3 carpels, rarely of 2, 4, or 5 carpels. Ovary sessile, often lobed, 

 sometimes appendaged ; styles distinct or united, slender throughout or 

 enlarged or dilated at the apex ; stigma usually minute, entire or lobed. Fruit 

 drupaceous, nut-like, capsular or a samara variously winged and sometimes also 

 crested . 



Receptacle high, usually pyramidal, usually 3-sided; fruit winged or 

 bristly, except in Aspicarpa and Brachypterys. 

 Fruit a samara, the wing reduced to a keel or beak in Brachypterys 

 and Aspicarpa. 

 Samaras with lateral distinct or united wings. 

 Androecium with 10 anther-bearing stamens. 



Samaras with a dorsal ridge, keel, or wing in addition to the 

 lateral wings. 

 Lateral samara-wings not lobed, sometimes united. 



Leaves with the stipules at the base of the petiole ; 



seeds with nearly equal cotyledons. 

 Leaves with the stipules on the petiole ; seeds with un- 

 equal cotyledons. 

 Lateral samara-wings lobed. 



Samaras 3-lobed, thus Y-shaped. 

 Samaras 4-lobed, thus X-shaped. 



Sepal-glands sessile, dorsally adnate. 

 Sepal-glands stalked, laterally adnate. 

 Samaras with thick cockscomb-like appendages between the 

 dorsal and lateral wings. 

 Androecium with 5 anther-bearing stamens, or fewer. 



Sepal-glands dorsally adnate ; normal flowers with 5 anther- 

 bearing stamens and 1 style, or with 3 stamens and 2 or 3 

 styles. 

 Sepal-glands basally adnate ; normal flowers with 5 anther- 

 bearing stamens and 3 styles. 

 Samaras with a single dorsal wing, this reduced to a keel or beak 

 in Brachypterys and Aspicarpa. 

 Androecium of lOperfector partially imperfect stamens ; styles3. 

 Stigma clavate or truncate. 



Stigma on the ventral edge of the dilated style-tips. 



Samara-wings thickened along the dorsal edge. 



Samara-wings thickened along the ventral edge. 



Androecium with 10 polleniferous anthers. 



Androecium with 4 polleniferous anthers. 



1 Some preliminary work on this family was accomplished several years ago by Charles 

 Budd Robinson, but this was interrupted by his appointment to the Philippines. 



Volume 25, Part 2, 1910] 117 



1. Mascagnia. 



2. HlRAEA. 



3. Triopteris. 



4. Tetrapteris. 



5. Adenoporces. 



6. Callaeum. 



7. Gaudichaudia. 



8. rosanthus. 



9. Banisteriopsis. 



10. Banisteria. 



11. Brachypterys. 



12. Stigmaphyllon. 



