Part 1, 1918] RUBIACEAE 5 
by longitudinal anterior slits. Disk depressed, explanate. Ovary 2-celled; style stout, 
filiform, glabrous, exserted, the 2 branches oblong, recurved, obtuse, papillose on the inner 
-side; ovules numerous, crowded on the placentae. Capsule turbinate, thick-coriaceous, 
areolate-truncate at the apex, 2-celled, loculicidally bivalvate from the apex, the valves re- 
curved. Seeds small, horizontal, obliquely subcuneate, the testa reticulate, the endosperm 
fleshy; embryo clavate, the cotyledons plano-convex, the radicle short, obtuse. 
Type species, Macrocnemum corymbosum R. & P. 
1. Condaminea corymbosa (R. & P.) DC. Prodr. 4: 402. 1830. 
Macrocnemum corymbosum R. & P. Fl. Per. 2: 48. 1799. 
Shrub or small tree, 2 meters high or more, with brown or yellowish-brown bark; branches 
stout, subterete, glabrous; stipules usually 4, sometimes connate into 2 or 3, 2—4.5 cm. long, 
oblong, acuminate or attenuate, puberulent at the base, glabrous above, glandular within at 
the base; petioles 1 cm. long or shorter, often wanting, very stout; leaf-blades obovate to 
oblong, 20-55 cm. long or more, 10—25 cm. wide, abruptly short-acuminate at the apex, gradually 
narrowed to the base or often contracted above the auriculate-cordate base, subcoriaceous, 
bright-green, glabrous, the costa stout and prominent beneath, the lateral veins 20-30 pairs, 
prominent, divergent, the veinlets also prominent; peduncles stout, 15-30 cm. long. glabrous or 
puberulent; bracts minute; pedicels short, stout; calyx campanulate, obtusely and irregularly 
5-dentate, about 8 mm. long, glabrous, green; corolla about 2.5 cm. long, purplish outside, 
green within; stamens 10-12 mm. long, the anthers 5 mm. long; capsule about 1.7 cm. long, 
bisulcate, glabrous, attenuate at the base, the valves 5-nerved; seeds about 4 mm. long, 
yellowish. 
TYPE LOCALITY: Peru. 
DISTRIBUTION: Panama to Peru. 
ILLUSTRATIONS: R. & P. Fl. Per. pl. 189; Mart. Fl. Bras. 65: pl. 124; E. & P. Nat. Pfl. 44: f. 6, H. 
2. CHIMARRHIS Jacq. Sel. Stirp. Am. 61. 1763. 
Trees, glabrous or pubescent. Leaves opposite, short-petiolate, the blades usually large. 
Stipules interpetiolar, convolute, caducous. Flowers small, arranged in axillary many-flowered 
long-pedunculate corymbose cymes, pedicellate, bracteate and bracteolate; hypanthium cam- 
panulate, the 5 lobes of the calyx very short, persistent; corolla short-funnelform, the tube very 
short, villous in the throat, the limb 5-lobate, the lobes recurved, valvate. Stamens 5, in- 
serted between the corolla-lobes; filaments filiform-subulate, exserted, pubescent at the base; 
anthers oblong, obtuse, dorsifixed, dehiscent by lateral slits. Disk orbicular. Ovary 2- 
celled; style short, thickened and sulcate above, shortly bifid, the lobes obtuse; ovules numerous, 
crowded on thick placentae peltately attached to the septum. Capsule small, bisulcate, 
septicidally bivalvate from the apex, the valves finally bipartite, the exocarp separating from 
the endocarp. Seeds numerous, very small, horizontal, crowded, angulate-globose or com- 
pressed, sometimes marginate, the testa reticulate; endosperm fleshy; embryo minute, bifid. 
Type species, Chimarrhis cymosa Jacq. 
Peduncles equaling or longer than the cymes; corolla-lobes obtuse. 1. C. cymosa. .- 
Peduncles much shorter than the cymes; corolla-lobes acute. 2. C. ferruginea. 
1. Chimarrhis cymosa Jacq. Sel. Stirp. Am. 61. 1763. 
Macrocnemum longifolium A. Rich. Mém. Soc. Hist. Nat. Paris 5: 279. 1834. 
Chimarrhis cymosa genuina Urban, Symb. Ant. 1: 410. 1899. 
Chimarrhis cymosa jamaicensis Urban, Symb. Ant. 1: 411. 1899. 
Chimarrhis cymosa microcarpa Urban, Symb. Ant. 1:411. 1899. 
Tree, sometimes 25 meters high; bark of the smaller branches brownish-gray; branchlets 
glabrous; stipules triangular-lanceolate, 2-4 cm. long, long-acuminate or long-attenuate, 
glabrous; petioles 1-3 cm. long, stout, semiterete, glabrous; leaf-blades obovate or oval-obovate, 
5-30 cm. long, 2-15 cm. wide, obtuse or acutish at the apex, sometimes abruptly acute, acute 
to attenuate at the base and often decurrent, thin, glabrous, or sometimes short-pilose beneath, 
paler beneath, the costa stout and prominent, the lateral veins slender, 7-16 on each side, 
