8 NORTH AMERICAN FLORA [VoLUME 32 
the lobes purple, 5-6 mm. long, ovate-oblong, obtuse, very thick, the throat puberulent; 
filaments very stout, the anthers oblong, 5 mm. long, equaling the corolla-lobes, opening by a 
small pore; capsule rounded-obovoid, 10 mm. long, 5—8 mm. wide, shallowly bisulcate, dark- 
brown; seeds brown, 1 mm. long or smaller. 
TYPE LOCALITY: Gorgona Island, Colombia. 
DISTRIBUTION: Costa Rica to Colombia; reported from Guatemala. 
5. PORTLANDIA P. Br. (Hist. Jam. 164; hyponym. 1756); L. Syst. 
- ed. 10.928. 1759. 
Gonianthes A. Rich. in Sagra, Hist. Cuba 11: 10. 1850. Not Gonianthes Blume, 1823. 
Shrubs or trees, glabrous or pubescent, often resinous. Leaves opposite or verticillate, 
petiolate or sessile, the blades large, usually coriaceous. Stipules intrapetiolar, connate with 
the petioles to form a sheath, usually persistent. Flowers large, axillary or terminal, the 
pedicels often bracteate; hypanthtum usually turbinate; calyx-lobes 4 or 5, short or elongate, 
persistent; corolla large, subcampanulate to funnelform or tubular-funnelform, the tube 5- 
angulate, glabrous in the throat, the limb 4- or 5-lobate, the lobes reduplicate-valvate or subim- 
bricate. Stamens 4 or 5, inserted at the base of the throat or at the base of the corolla-tube, 
the filaments filiform, pubescent; anthers basifixed, linear, included or short-exserted. Disk 
lobate. Ovary 2-celled; style filiform, the apex entire, bilobate, or with 2 long branches; 
ovules numerous, crowded on swollen placentae longitudinally adnate to the septum. Capsule 
thick-coriaceous, terete, costate, or angulate, loculicidally bivalvate from the apex. Seeds 
numerous, usually compressed, angulate, the testa usually granulate; endosperm fleshy; 
embryo minute, the cotyledons ovate, the radicle terete. 
Type species, Portlandia grandiflora L. 
Leaves sessile or nearly so, the blades cordate or subcordate at the base. 
Capsule about 2 cm. long. 1. P. Harrisii. 
Capsule 1 cm. long or less. 
Flowers pedicellate; leaf-blades elliptic, 5-10 cm. long; corolla pink- 
ish. 22 PA nibens: 
Flowers sessile. 
Calyx-lobes linear, about as long as the hypanthium; leaf-blades 
1.5—3 cm. long, nearly orbicular. 3. P. sessilifolia. 
Calyx-lobes triangular, shorter than the hypanthium ; leaf-blades 
4-11 cm. long, oval-oblong. 4. P. Shaferi. 
Leaves petiolate, narrowed or rounded at the base. 
Leaf-blades rounded or very obtuse at the apex; branchlets usually 
puberulent. 
Flowers surrounded by an involucre of numerous bractlets, 
terminal. 5. P. involucrata. 
Flowers exinvolucrate, often axillary. 
Leaf-blades 1.5—3 cm. long. 6. P. uliginosa. 
Leaf-blades mostly 3.5—8 cm. long. 7. P. elliptica. 
Leaf-blades acute or acuminate at the apex; branchlets glabrous. 
Lobes of the calyx and corolla 4 each. 8. P. Ghiesbreghtiana. 
Lobes of the calyx and corolla 5 each. 
Calyx-lobes oblong to ovate, 20-35 mm. long. 9. P. grandiflora. 
Calyx-lobes linear or subulate, or in one species triangular, but 
then only 1-6 mm. long. 
Capsule 1.2 cm. long or less, shorter than the calyx-lobes; 
corolla about 5 cm. long. 10. P. pendula. 
Capsule more than 1.2 cm. long, longer than the calyx-lobes. 
Capsule only obscurely angled. 
Corolla 20-25 cm. long; leaf-blades’ 12-20 cm. long, 
the margins plane. 11. P. Lindeniana. 
Corolla 5-8 cm. long; calyx-lobes linear; leaf-blades 
3.5-14 cm. long. 
Margins of the leaves plane; stipules not cuspidate. 
Calyx-lobes 7-15 mm. long; corolla-lobes 
triangular. 
Corolla crimson. 12. P. coccinea. 
Corolla white. 13. P. albiflora. 
Calyx-lobes 1-6 mm. long; corolla-lobes 
semi-orbicular. 14. P. microsepala. 
Margins of the leaves revolute; stipules long- 
cuspidate. 15. P. mexicana. 
Capsule acutely angled. 
