ParT 1, 1918] RUBIACEAE 19 
lobes obtuse, valvate. Stamens 4, inserted in the throat of the corolla; filaments short; anthers 
dorsifixed, usually exserted. Disk small. Ovary 2-celled; style slender, the 2 branches short, 
linear, obtuse; ovules numerous, rarely few, inserted on placentae attached to the base or rarely 
to the middle of the septum. Capsule small, usually membranaceous, terete or angulate, 
globose or turbinate, wholly attached to the hypanthium, loculicidally dehiscent at the apex or 
through its whole length, few-seeded. Seeds angulate or subglobose, the testa smooth or 
minutely granulate; endosperm fleshy; embryo clavate. 
Type species, Oldenlandia corymbosa L. 
Flowers in few-flowered cymes, usually long-pedicellate. 
Corolla inconspicuous, about 1 mm. long; plants annual. 1. O. corymbosa. 
Corolla conspicuous, 3 mm. long or longer; plants annual or perennial. 
Corolla 3mm. long; flowers partly sessile in the forks of the branches; 
plants annual. 2. O. Greenei. 
Corolla 5-8 mm. long; flowers all pedicellate; plants perennial. 
Leaf-blades lanceolate to broadly ovate, petiolate. 3. O. microtheca. 
Leaf-blades linear, sessile. 
Leaves 4-12 mm. long, succulent; corolla abruptly dilated into 
the limb, 5-6 mm. long. 4. O. Pringlei. 
Leaves 15-45 mm. long, not succulent; corolla gradually 
dilated from base to throat, 8 mm. long. 5. O. xestosperma. 
Flowers solitary on slender axillary or terminal pedicels, or crowded in dense 
axillary fascicles. 
Pedicels shorter than the calyx. 
Calyx-lobes eciliolate; plants perennial; capsule granulate. 6. O. Boscii. 
Calyx-lobes ciliolate; plants annual; capsule smooth. 
Stems white-hirsutulous; leaves short-petiolate. 7. O. uniflora. 
Stems glabrous; leaves sessile or nearly so. 8. O. fasciculata. 
Pedicels much longer than the calyx. 
Corolla equaling or shorter than the calyx-lobes. 
Stems scaberulous; leaf-blades oval or elliptic. 9. O. crystallina. 
Stems glabrous; leaf-blades linear or linear-lanceolate. 10. O. herbacea. 
Corolla much longer than the calyx-lobes. 
Leaf-blades suborbicular, as broad as long; capsule turbinate, 
longer than broad. 11. O. callitrichoides. 
Leaf-blades elliptic to rounded-ovate, longer than broad; capsule 
subglobose, as broad as long. 
Leaf-blades mostly ovate-deltoid, long-petiolate, broadly 
rounded or truncate at the base. 12. O. ovata. 
Leaf-blades mostly elliptic, sessile or short-petiolate, acute or 
obtuse at the base. 13. O. capillipes. 
1. Oldenlandia corymbosa L. Sp. Pl. 119. 1753. 
Hedyotis corymbosa Lam. Tab. Encye. 1: 272. 1791. 
Gerontogea corymbosa Cham. & Schlecht. Linnaea 4: 154. 1829. 
Annual, erect or decumbent, glabrous except upon, the leaf-blades, usually much branched, 
the branches slender or stout, 1-4 dm. long, obtusely quadrangular; stipules 1-1.5 mm. long, 
whitish, truncate, bearing 1 or more long bristles; leaves sessile or subsessile, the blades linear 
to linear-lanceolate, 0.2-3.5 cm. long, 0.5—5 mm. wide, acute to attenuate at the apex, acute to 
attenuate at the base, herbaceous, deep-green above and usually scaberulous, pale and glabrous 
beneath, l-nerved, the margins scaberulous, often subrevolute; flowers in axillary cymes, or 
rarely solitary, the cymes usually 3-flowered, the peduncles filiform, 5-15 mm. long, often 
recurved, the pedicels longer or shorter than the peduncles; hypanthium less than 1 mm. 
long, the calyx-lobes usually longer, triangular, acuminate; corolla white, short-salverform, 
equaling or slightly exceeding the calyx-lobes; capsule subglobose, 1.5—-2 mm. broad and almost 
as long, bisulcate; seeds 0.2-0.3 mm. long, brown. 
TYPE LOCALITY: Tropical America. 
DISTRIBUTION: Costa Rica and Panama; Jamaica and the Lesser Antilles; also in South America 
and in tropical regions of the Old World. 
ILLUSTRATIONS: Gaertn. Fruct. pl. 30, f. 3; Descourt. Fl. Ant. pl. 50; Plum. Ic. Burm. pl. 2/2, 
jf. 1; Lam. Tab. Encye. pl. 61, f. 4. 
2. Oldenlandia Greenei A. Gray, Proc. Am. Acad. 19: 77. 1883. 
Slender erect annual, 5—30 cm. high, simple or sparsely branched, glabrous throughout, 
the internodes few, equaling or shorter than the leaves; stipules minute, triangular, cuspidate; 
leaves sessile or the lower short-petiolate, the blades linear or spatulate-linear, or those of the 
