Part 2, 1921] RUBIACEAE 95 



Anthers borne on long filaments. 



Corolla symmetric, the tube very slender, terete, the limb 



4-5-lobate. 38. ExosTBMA. 



Corolla often asymmetric, the tube obconic or gibbous, the 



limb 6-8-lobate. 39. Coutarea. 



Anthers sessile or on very short filaments. 



Anthers sessile. 40. Blepharidium. 



Anthers on short filaments. 35. Ravnia. 



29. LADENBERGIA Klotzsch, in Hayne, Arzn. Gew. 14: pi. 15. 



1846. 

 Cascarilla Wedd. Ann. Sci. Nat. III. 10: 10. 1848. Not Cascarilla Raf. 1838. 



Trees or shrubs, the branchlets terete or angulate. Leaves opposite or verticillate, peti- 

 olate, coriaceous. Stipules interpetiolar, deciduous, glandular within at the base. Flowers 

 small or of medium size, white, arranged in terminal panicles; hypanthium ovoid or turbinate, 

 pubescent; calyx cupular, 5-lobate, valvate, persistent or deciduous; corolla subfunnelform or 

 salverform, pubescent outside, the tube glabrous within, the limb 5-lobate, valvate, the lobes 

 papillose within and on the margins. Stamens 5, inserted below the middle of the tube; fila- 

 ments short; anthers linear, dorsifixed, included. Disk pulvinate. Ovary 2-celled; style 

 filiform, the branches oblong or linear, included; ovules numerous, imbricate, the placentae 

 terete, adnate to the septum. Capsule oblong, clavate, or cylindric, 2-celled, septicidally 

 bivalvate from the apex to the base. Seeds numerous, imbricate, peltate, the body oblong, 

 surrounded by a membranaceous, dentate or lacerate wing, the placentae terete or angulate; 

 endosperm fleshy; cotyledons ovate; radicle terete. 



Type species, Ladenbergia undata Klotzsch. 



1. Ladenbergia undata Klotzsch, in Hayne, Arzn. Gew. 14: pi. 15. 



1846. 



Cascarilla undata Wedd. Ann. Sci. Nat. III. 10: 12. 1848. 



Shrub or tree, the branchlets stout, obtusely angulate, pilose when young; stipules ovate, 



about 3 cm. long, obtuse or acutish, coriaceous, the margins revolute; petioles stout, 1-3.5 cm. 



long, pilose or glabrate; leaf-blades oblong or oval-oblong, 7-20 cm. long, 3.5-10 cm. wide, 



rounded to acutish at the base, obtuse or acutish at the apex, thick-coriaceous, more or less 



bullate, lustrous above, glabrous, the venation often impressed, brownish beneath, pilose, 



especially along the veins, the lateral nerves very prominent, about 10 on each side, the margin 



strongly revolute; inflorescence cymose-paniculate, pedunculate, few-flowered, the flowers 



sessile or short-pedicellate; hypanthium densely ferrugino-pilose with mostly appressed hairs; 



calyx-lobes ovate-triangular, acute; corolla about 3.5 cm. long, densely sericeous-pilose with 



ferruginous hairs, the lobes lance-oblong, acute, about half as long as the tube; capsule linear- 



clavate, about 4.5 cm. long and 6 cm. thick, brownish; seeds reddish-brown, broadly winged, 



the wing lacerate-dentate. 



Type locality: Near Merida, Venezuela. 

 Distribution: Costa Rica to Colombia and Venezuela. 



30. ALSEIS Schott; Spreng. Syst. 4 2 : 404. 1827. 



Trees or shrubs, more or less pubescent, with terete branches. Leaves opposite, petiolate, 

 the blades of medium size. Stipules interpetiolar, short or elongate. Flowers small, white 

 or yellow, ebracteolate, spicate, the spikes simple or branched, axillary and terminal; hypan- 

 thium obconic; calyx 5-lobate, the lobes broad or narrow, deciduous; corolla cylindric, short, 

 pilose within, the limb 5-lobate, the lobes valvate. Stamens 5, inserted at the base of the 

 corolla- tube ; filaments elongate, villous, complanate at the base; anthers versatile, oblong, 

 exserted, bipartite at the base. Disk broad, swollen. Ovary 2-celled; style elongate, exserted, 

 pilose or glabrous, bipartite, the branches recurved, obtuse; ovules numerous, imbricate, the 

 placentae elongate, pendulous from the apex of the cell. Capsule oblong-turbinate, 2-celled, 

 septicidally bivalvate from the apex, the valves entire, crustaceous. Seeds linear-fusiform, 

 the body oblong, the testa thick, long-produced at each end; endosperm fleshy; embryo elon- 

 gate, the cotyledons narrow, the radicle terete, subelongate. 



Type species, Alseis floribunda Schott. 



