Part 2, 1921] RUBIACEAE 



thium obovoid; calyx large, with a distinct tube, 4-lobate, the lobes very l)road, imbricate, 

 ciliolate; corolla salverform, coriaceous, the tube slender, elongate, densely villous within 

 except near the base, the limb 4-lobate, the lobes broad, spreading, imbricate in bud, one of 

 the lobes exterior. Stamens 4, inserted in the throat of the corolla; anthers sessile, dorsi- 

 fixed below the middle, linear, obtuse, included. Disk annular. Ovary 2-celled; style filiform, 

 included, the stigma bilobate, the lobes linear, elongate, acute; ovules numerous, imbricate, 

 winged, the placentae attached to the septum. 



Type species, Blepharidium guatemalense Standley. 



1. Blepharidium guatemalense Standley, Jour. Wash. Acad. 

 8:59. 1918. 



Branchlets stout, green, glabrous, sparsely whitish-lenticellate, the internodes elongate; 

 stipules ovate-triangular, 2-2.5 cm. long, acuminate or cuspidate-acuminate, brown, glabrous 

 outside, within sericeous-pilose at the base and bearing numerous glands; petioles stout, 

 2.5-5 cm. long, glabrous; leaf-blades oval-oblong, 17-30 cm. long, 8-14.5 cm. wide, obtuse or 

 acute at the base, obtuse or acute at the apex, concolorous, glabrous above, the costa impressed, 

 the other venation prominulous, sparsely short-pilose beneath along the prominent costa, 

 the lateral veins prominent, about 8 on each side, arcuate-ascending; peduncles 3-flowered, 

 slender, 7-21 cm. long, glabrous, the pedicels stout, 0.3-2 cm. long; bracts foliaceous, oval, 

 about 1 cm. long, the bractlets broadly ovate, 3-4 mm. long, deciduous; hypanthium glabrous, 

 4—5 mm. long; calyx 4—5 mm. long, 7-8 mm. broad, the lobes half as long as the tube, broader 

 than long, rounded or truncate, minutely ciliolate; corolla-tube about 6 cm. long, 4-5 mm. 

 thick, glabrous outside, the lobes oval, about 1 cm. long; anthers 7 mm. long; stigma-lobes 

 6-7 mm. long. 



Type locality: Forest along Saklak River, below Secanquim, Alta Verapaz, Guatemala, alti- 

 tude 300 meters. 



Distribution: Known only from the type locality. 



V. NAUCLEEAE. Shrubs or trees, sometimes scandent. Flowers per- 

 fect, numerous, crowded on a globose receptacle; corolla funnelform, with 

 an elongate tube, the lobes short, imbricate or valvate. Anthers short, 

 usually subsessile. Ovary 2-celled; stigma clavate, capitate, or fusiform; 

 ovules numerous or solitary. Fruit a many-celled syncarp or a capsule or 

 dicoccous. Seeds minute, winged or exalate; radicle superior. 



Ovules solitary; plants erect, unarmed. 41. Cephalanthus. 



Ovules numerous; plants climbing by recurved spines. 42. Ourouparia. 



41. CEPHALANTHUS L. Sp. PI. 95. 1753. 



Acrodryon Spreng. Syst. 1: 386. 1825. 

 Silamnus Raf. Sylva Tell. 60. 1838. 

 Axolus Raf. SylvaTell. 61. 1838. 

 Gilipus Raf. Sylva Tell. 61. 1838. 

 Eresimus Raf. Sylva Tell. 61. 1838. 



Erect shrubs or trees, glabrous or pubescent, the branchlets terete or obtusely quadran- 

 gular. Leaves opposite or 3-4-verticillate, petiolate. Stipules intrapetiolar, short. Flowers 

 small, white or yellow, sessile in dense globose heads, intermixed with setaceous or paleaceous 

 bractlets; hypanthium turbinate; calyx short-tubular, unequally 4-5-dentate or 4-lobate, 

 often glanduliferous; corolla tubular-funnelform, the throat glabrous or pilose, the limb 4-5- 

 lobate, the lobes erect or spreading, imbricate, one exterior, often with glands in the sinuses. 

 Stamens 4, inserted in the throat of the corolla; filaments short; anthers dorsifixed, oblong, 

 bicuspidate at the base. Disk inconspicuous. Ovary 2-celled; style filiform, elongate; 

 stigma capitate or clavate; ovules solitary, pendulous from the top of the cell, anatropous, the 

 funicle thickened. Fruit turbinate, 2-4-celled, the cells indehiscent, 1-seeded, or one or more 

 of them empty. Seeds oblong, pendulous, covered at the apex with a white aril, the testa 

 membranaceous, sometimes short-winged; endosperm subcartilaginous ; cotyledons plane, 

 linear-oblong, the radicle slender, superior. 



Type species, Cephalanthus occidentalis L. 



