Part 2. 1921] RUBIACEAE 135 



subulate; hypauthium and calyx strigose, 1.5 mm. long, the 4 calyx-lobes triangular, acute, 



shorter than the hypanthium; corolla white, 7-8 mm. long, strigose, the tube stout, ihe 4 



lobes ovate, about one third as long as the tube; style bifid at the apex; fruit subglobose, 



fleshy at maturity, dicoccous, 2-3 mm. broad; seeds brown, lustrous. 



Type locality: Cuba. 

 Distribution: Cuba. 



3. Duggena spicata (Lam.) Standley, Contr. U. S. Nat. Herb. 

 18: 126. 1916. 



Lygistum s pic alum Lam. Tab. Encyc. 1: 286. 1791. 



Duggena Richardii West, Bidr. St. Croix 269. 1793. 



Pelesia spicata Sw. Fl. Ind. Occ. 1945, excluding description. 1806. 



Coccocipsilum spicalum H. B. K. Nov. Gen. & Sp. 3: 406. 1820. 



Hedyolis secunda Spreng.; Schultes, in R. & S. Syst. Veg. Mant. 3: 133. 1827. 



Gonzalagunia Coccocypselum Cham. & Schlecht. Linnaea 4: 196. 1829. 



Gonzalea spicata DC. Prodr. 4: 437. 1830. 



Gonzalea spicata glabrata DC. Prodr. 4: 437. 1830. 



Gonzalea Coccocypselum Steud. Nom. ed. 2. 1: 701. 1840. 



Gonzalagunia hirsuta K. Schumann, in Mart. Fl. Bras. 6 6 : 291. 1889. Not Justicia hirsuta Jacq. 1760. 



Gonzalagunia hirsuta dicocca K. Schumann, in Mart. Fl. Bras. 6 6 : 292, in part. 1889. 



Gonzalagunia hirsuta f. leucocarpa Kuntze, Rev. Gen. 284. 1891. 



Gonzalagunia hirsuta f. cyanocarpa Kuntze, Rev. Gen. 284. 1891. 



Gonzalea grisea Wernham, Jour. Bot. 51: 219. 1913. 



Duggena grisea Standley, Contr. U. S. Nat. Herb. 18: 125. 1916. 



Shrub or small tree, 0.5-4 meters high, the branches slender, densely sericeous-strigose 

 when young; stipules 4-11 mm. long, long-subulate from a triangular base; petioles 0.2-2.5 

 cm. long; leaf-blades broadly ovate, oblong-ovate, elliptic-oblong, or lanceolate, 6.5-19 cm. 

 long, 2-8.5 cm. wide, acute to rounded at the base, often abruptly decurrent, usually sub- 

 abruptly long-acuminate or attenuate at the apex, membranaceous, bright-green above, 

 appressed-pilose or glabrate, the venation plane or prominulous, paler beneath, appressed- 

 pilose, especially along the veins, the lateral veins prominulous, numerous, arcuate, usually 

 ascending at an angle of 50° or less; inflorescence spiciform, 8-40 cm. long, short-pedunculate, 

 often dense, secund, the cymules 1-few-flowered, sessile or short-pedunculate, the pedicels 

 2 mm. long or shorter, the bracts linear or subulate, sometimes foliaceous; calyx and hypan- 

 thium densely appressed-pilose, the hypanthium 1.5-2 mm. long, the 4 calyx-lobes equaling 

 or longer than the tube, linear, lanceolate, oblanceolate, or oblong, acute; corolla white, 10- 

 15 mm. long, densely appressed-pilose outside with white, often long hairs, the tube stout, 

 ampliate above, the 4 lobes rounded, half as long as the tube or shorter, the throat densely 

 white-pilose; stamens 4; style bifid at the apex; fruit subglobose, fleshy at maturity, white 

 or blue, dicoccous, 3-4 mm. broad; seeds dark-brown, lustrous. 



Type locality: Antilles. 



Distribution: Porto Rico, Hispaniola, and the Lesser Antilles; also from Venezuela and the 

 Guianas to Brazil; reported from Veracruz and Costa Rica. 



4. Duggena rudis Standley, Contr. U. S. Nat. Herb. 18: 

 125. 1916. 



Branches very slender, brownish, densely whitish-strigose; stipules subulate, 3-6 mm. 

 long; leaves subsessile, the stout petioles 2 mm. long or shorter, the blades ovate or ovate- 

 oblong, 3.5-7 cm. long, 1.4-2.5 cm. wide, rounded at the base, subabruptly long-acuminate at 

 the apex or acute, membranaceous, bright-green above, copiously strigose and strigillose, the 

 venation prominulous, paler beneath, densely whitish-strigose when young, glabrate in age 

 except along the veins, the lateral veins prominent, 5 or 6 on each side, arcuate-ascending at an 

 angle of 45° or less; inflorescence interrupted-spiciform, 6-9 cm. long, slender, short-peduncu- 

 late, the cymules 1-3-flowered, sessile or subsessile, the pedicels 1 mm. long or shorter; bracts 

 linear-subulate, 3 mm. long or shorter; calyx and hypanthium very densely setose-pilose, the 

 4 calyx-lobes linear-attenuate, 1-2 mm. long; corolla white, 5 mm. long, densely appressed- 

 pilose, the tube stout, the lobes one third as long as the tube or shorter; stamens 4; style 

 quadrifid; fruit tetracoccous, depressed-globose, 2.5 mm. broad, densely setose-pilose with 

 subappressed hairs; seeds brown. 



Type locality: In shady places along the Chagres River, near El Vigia, Panama. 



Distribution: Panama and Colombia. 



