Part 2, 1921] RUBIACEAE 137 



8. Duggena panamensis (Cav.) Standley, Contr. U. S. Nat. 

 Herb. 18: 126. 1916. 



Buena panamensis Cav. Anal. Hist. Nat. 2: 279. 1800. 



Gonzalea eornifolia H. B. K.. Nov. Gen. & Sp. 3: 325. 1819. 



Gonzalca panamensis Spreng. Syst. 1: 417. 1825. 



Gonzalea leptantha A. Rich, in Sagra, Hist. Cuba 11: 16. 1850. 



Gonzalagunia panamensis K. Schumann, in Mart. I'M. Bras. 6 6 : 292. 1889. 



Gonzalagunia leptantha Maza. Anal. Soc. Ksp. Hist. Nat. 23: 289. 1894. 



Gonzalea Hayesii Wcrnham, Jour. Hot. 51: 219. 1913. 



Duggena Hayesii Standley, Contr. U. S. Nat. Herb. 18: 125. 1916. 



Duggena leptanlha Standley, Contr. U. S. Nat. Herb. 18: 125. 1916. 



Shrub, 0.6-3 meters high, the branches slender, brownish, densely sericeous or strigosc 

 when young; stipules triangular, subulate-acuminate, 3-8 mm. long; petioles 0.3-2.2 cm. long; 

 leaf-blades ovate, ovate-oblong, lance-oblong, or lanceolate, 7-14 cm. long, 1.8-6 cm. wide, 

 usually acute or obtuse at the base, rarely rounded, gradually or abruptly acuminate or long- 

 attenuate at the apex, membranaceous, bright-green above, densely or sparsely strigillose or 

 scaberulous or glabrate, the venation plane or prominulous, slightly paler beneath, sparsely 

 or densely setose-pilose or soft-pilose or glabrate, the lateral veins prominent, 5-9 on each side, 

 arcuate, usually ascending at an angle of 50° or less; inflorescence spiciform, usually dense, 

 8-24 cm. long, short-pedunculate, the cymules 1- few-flowered, sessile or subsessile, the pedicels 

 2 mm. long or shorter, the bracts subulate, 2-3 mm. long; calyx and hypanthium 1.5-2 mm. 

 long, strigillose or short-pilose or glabrate, the 4 calyx-lobes deltoid, obtuse or acute, shorter 

 than the tube; corolla white, 10-17 mm. long, the tube very slender, glabrous or sparsely 

 pilose, the 4 lobes 2-2.5 mm. long, usually pilose outside, sparsely tomentose within; stamens 4, 

 included; style quadrifid; fruit globose, tetracoccous, 3-4 mm. in diameter, fleshy at maturity; 

 seeds dark reddish-brown. 



Type locality: Ancon Hill, Panama. 



Distribution: Veracruz to Michoacan and Colombia; Cuba and the Isle of Pines. 



Illustration: Cav. Ic. pi. 571. 



Doubtful or excluded species 



Duggena nivea (Bartl.) Standley, Contr. U. S. Nat. Herb. 18: 125. 1916. {Gonzalea 

 nivea Bartl.; DC. Prodr. 4: 436. 1830. Conzalagunia nivea Kuntze, Rev. Gen. 284. 1891.) 

 Described from Mexico. Probably a species of Rondcletia. 



Gonzalagunia parviflora Schlecht. & Cham. Linnaea 6: 414. 1831. {Gonzalea 

 parviflora G. Don, Gen. Hist. 3: 537. 1834.) Described from Mexico. The description is 

 too brief for determination. 



Gonzalea glabra S. Wats. Proc. Am. Acad. 25: 152. 1890. This is Cilharexylum 

 glabrum (S. Wats.) Greenman. 



Gonzalea secunda Mart. & Gal. Bull. Acad. Brux. 11': 235. 1844. {Gonzalagunia 

 secunda Kuntze, Rev. Gen. 284. 1891.) Described from Jalapa, Veracruz. Perhaps a 

 synonym of Duggena panamensis. 



46. CASSUPA Humb. & Bonpl. PI. Aequin. 1: 43. 1808. 



Small trees, pubescent, the branchlcts thick, terete or subangulate. Leaves opposite, 

 petiolate, large, coriaceous. Stipules interpetiolar, acuminate, persistent. Flowers large, 

 red or white, cymose, thyrsiform-paniculate, sessile or pedicellate, bracteate and bracteolate; 

 hypanthium subglobose ; calyx short, obscurely 6-lobate, persistent ; corolla coriaceous, funnel- 

 form, the tube elongate, often tubcrculate, glabrous within, the throat villous, the limb 6- 

 lobate, the lobes short, valvate, villous within. Stamens 6, inserted in the throat of the 

 corolla; filaments subulate; anthers dorsifixed, acuminate, included. Disk cupular. Ovary 

 2-celled; style slender, thickened at the apex, with 2 short rounded lobes; ovules numerous, 

 the placentae swollen, adnate to the septum. Fruit baccate, globose, 2-celled. Seeds nu- 

 merous, minute, angulate. 



Type species, Cassapa verrucosa Humb. & Bonpl. 



