105 157 
Subgenus 5. Glaphyropteris (Presl) C. Chr. 
Biolog. Arbejder tilegnede Eug. Warming p. 80. 1911. 
Glaphyropteris Presl, Abhandl. bóhm. Ges. Wiss. V. 5: 344, 1848. 
A small subgenus of mostly large species with a bipinnatifid lamina and an 
often very large (1 cm or more long), acute aérophore at the bases of the pinnz 
beneath and similar but smaller aérophores at the bases of the midribs 
of the segments (not found in D. mapiriensis) '- 
Segments close, rectangular with broad, bluntly rounded apex and entire 
margins, rarely the apex is acute. Veins very close and numerous, simple, the 
basal ones reaching the margins above sinus. Pubescence somewhat variable (see 
below), the under-surface of most species covered with numerous, sessile, red 
glands, which are deciduous and therefore not found in older specimens. 
By these characters the six species referred to Glaphyropteris differ from 
Lastrea, but I fear that the characters mentioned are not sufficient for the segre- 
gration of the two proposed subgenera. If one should prefer to treat Lastrea as a 
genus, which would be a very natural treatment, Glaphyropteris ought to be referred 
to it as a subgenus. It is true that the typical species of Glaphyropteris, D. decus- 
sata, is very different from all species referred to Lasírea, but it is no doubt 
intimately related to the three first species mentioned below. These three species 
show both the characters of Lastrea and Glaphyropteris besides some others peculiar 
to them alone; with the same right they could be referred to Lastrea, to a proper 
subgenus or to Glaphyropteris. I prefer here to refer them to a proper section of 
Glaphyropteris, which they are perhaps nearest related to. The subgenus thus is 
divided into two groups each including three species: 
1. Group of D. Thomsonii. The three species belonging here resemble species 
of Lastrea, especially those related to D. rudis, by the shape of the lamina, which 
is abruptly narrowed downwards with 3—4 pairs of glanduliform warts along the 
stipe, and further by the shortly and antrorsely setose costze above; the veins are 
not so close as in the species af Glaphyropteris proper, which they resemble by 
the presence of aérophores at the base of the midribs beneath. 
The pubescence of the coste beneath (partly also of the rachis) is peculiar 
and different from all species of Lastrea and Eu-Glaphyropteris. The coste be- 
neath are, namely, shortly and often densely cinereo-tomentose by sessile, 2—3- 
branched hairs, which do not resemble the stalked, branched hairs of Goniop- 
teris. Further the sori seem to be indusiate at least in D. Canadasii and D. 
macradenia. ‘The three species belonging here were in my former papers referred 
to the group of D. opposita. (’Revision” nr. 67, 68 and 79). 
! The most important function of the aérophores or better "pneumatophores" appears to be 
during the development of the leaf, while the growing parts of this are covered by mucilage. Very 
likely they are provisions for the aeration of the young parts; in the developed leaf they are shriveled. 
See Bower, Annals of Botany 24: 427—428, foot-note, 1910. 
D. K. D. Vidensk. Selsk, Skr., 7. Rekke, naturvidensk, og mathem, Afd. X. 2. 21 
