106 MR. G. BUSK ON THE ANCIENT OR 
As it was found in the dark cave-earth above the stalagmite floor of the Genista cave, 
in which one of the astragali was also discovered, it may be probable that the entire 
fourth metatarsal belonged to the same foot with that bone, although the depths at 
which they occurred were about 7 feet apart. 
1. The entire third metatarsal (Pl. XVI. figs. 1-5) is 6-7 long; the proximal end 
measures 1’-6 x 1’-9, and the distal the same. The transverse diameter of the shaft at 
the middle is 1-8, and at the smallest part 1’°6; the least circumference is 42, and 
the perimetral index ‘626. The corresponding dimensions in a perfect third metatarsal 
in the Brady collection (of which an outline figure is given in Pl. XI. fig. 2) are 
almost identically the same—viz. length 6’°7, proximal end 1"-7 x 1'"9, and the distal 
1"-6x1"-9. The width of the shaft at the middle is 1/8, and at the smallest part 1’"6, 
the least circumference 4"°3, and the perimetral index *642. Nor do the bones present 
any appreciable difference whatever in conformation. 
The corresponding measurements in 2. megarhinus, No. 19842, B. M., from Grays, 
are—length 8"-0, proximal end 2!5 x —?, distal 1-9 x 28, transverse diameter of shaft 
at middle 2:3, and at the smallest part 2’:1, the least circumference 5':2, and perimetral 
index 650. 
The following Table shows the relative dimensions of the third metatarsal in other 
instances of fossil and recent Rhinoceros; but the above will suffice to show the close 
resemblance between the Gibraltar metatarsal and that of R. hemitachus, and at the 
same time its divergence from the megarhine type :— 






a=) 
S zr) Ss g oS 
roe 3 3 5 gy | U3 
He 5 es & os Sc 
Dimensions of third metatarsal in yarious se a 3 3 a q E aa ga 
species of Rhinoceros. 5 a g S as a £ aS 3 
Sse see fost e Wek ee 
3 a as aa 8 'S 24 
=) = <4 = BH Ay Ay 
Guibraltians sp... 1... cof sisisee teeters siete 66 |16 x1:9)1'55x1:9)'8 x1:75) 42 | -636 | -218 
R. hemiteechus, Uford, No. 20245, B.M. ..| 68 {16 x2-2) x2-0)/85x1:8 | 44 | -647 | -211 
A Pr Grays, No. 20267, B.M...| 6-4 |1:7 x1:9/1'6 x1:9'8 x16 | 3-9 | -609 | -200 
= Rs Sir A. Brady’s collection ..| 6°7 |1°6 x1-9/1°55x1:9/9 x1:8 4-2 | -621 | -200 
” ” ” eine Sova eee. 6:3 16 x 1-9 14 x18 
» megarhinus, Grays, No. 19842, B.M. ..| 8:0 x2-5/1-9 x 2:39 ~ 21 5:2 | -650 | -233 
i AA No. 23761, g. B.M....... 7-75 |1:9 x 2-4|1-85 x 2°39 x2:0 | 5:0 | -657 | -222 
» tichorhinus, Kent’s Cavern .......... 66 1:75 x 2-2 x 2°2/1:0 x 2-0 5:0 | -757 | -200 
», etruscus, Val d’Arno, No. 28808, B.M...| 6°85 |1°8 x2-0/1:5 x1-7/9 x16 | 3:9 | -567 | -180 
» meisivus, No. 29646, B.M. .......... 62 |1-7 x2-0)1'8 x2:0/9 x16) 41 | -668 | -180 
BI ULCOTI S| (CLUaMELLOG)) «tas see was sfeis oe 5:9 {16 x1-911:'5 x1-9'8 x1-65) 4:2 | -712 | -206 
Fr SUMATENSIE RECISo els cae eene ea 6:3 x 2-0 x 2:0 
saun CN TCOTTLIS MEL: OS eee co hee Ate ts, 79 |2:2 x2-51-9 x2-8/2:2x25 









2. The fourth metatarsal, represented in Pl. XVII. (figs. 1-5), is quite perfect, and 
of a fully mature animal, inasmuch as there is no trace whatever of the epiphysial 
suture. 
