Phyllophora Brodiæi and Actinococcus subcutaneus. 30 
plants do not exist. The antheridia and the procarps usu- 
ally occur on the same individual; they arise either in 
particular sexual folioles or in the upper margin of the flat 
fronds. The antheridia quite agree with those in Phyll. 
membranifolia. As in this species the procarps consist of 
a three-celled carpogonial branch and a bearing cell or 
auxiliary cell, but the carpogonial branch is rather variable, 
perhaps a consequence of degeneration, and it is probable 
that fertilization does not take place; at all events it has 
not been ascertained. The auxiliary cell much resembles 
that in Ph. membranifolia; in both species it has first one 
nucleus but later becomes plurinuclear. But while the latter 
is a typical diplobiontic species in which the auxiliary cell 
pushes out a number of outgrowths developing into the 
gonimoblast filaments, the auxiliary cell in Ph. Brodiei 
likewise pushes out a number of protuberances, but these 
give rise to cell-filaments forcing their way between the 
cells of the gametophyte in various directions but at first 
especially outwards where they give rise to a wartlike 
excrescence which develops into a nemathecium in the inner 
part of which the enlarged auxiliary cell or central cell is to 
be seen. The radiating filaments of the nemathecium give 
rise to seriate tetrasporangia, which ripen in winter. The 
system of cell-filaments issuing from the auxiliary cell in 
Ph. Brodiæwi and their products thus represent the tetra- 
sporophyte which is not here as in other Florideæ an inde- 
pendent free-living organism, but grows out in continuity 
with and “parasitically” upon the gametophyte generation. 
Cystocarps are never produced, the carposporophytic phase 
has been abandoned, and in its place a tetrasporophyte is 
developed. The vegetative part of the latter is only repre- 
sented by the intramatrical cell-filaments. 
3* 
