230 DR. R. W. SHTJFELDT ON [Apr. 1, 



represented by an inconspicuous intercalated bit of tbin cartilage, 

 barely preventing tbe contact of the tibio-fibulare with the first two 

 metfitarsals. Proxinnal epiphyses of these latter, however, are 

 moulded to meet the ends of perfect articulation. Subcuboidal in 

 form, tarsale 3 is a well-ossified bonelet articulating with second and 

 third metatarsals, with tarsale 2, and with the tibio-fibnlare, or the 

 co-ossified mass representing the elements of tbe proximal row. The 

 basal ends of the 2nd and 3rd metatarsals are markedly smaller than 

 they are in the 1st and 4rtb, indeed in tbe latter it has its proximal 

 extremity very conspicuously expanded. Tarsale 4 is a larger 

 nodule of bone that articulates with the tarsal elements upon 

 either side of it, with tbe 4th metatarsal, and with the tibio-fibulare. 

 Finally, more remarkable than any of the rest is tarsale fi ; it is a 

 wonderfully irregular bone in form, and peculiar in other respects. 

 It articulates by merely a vertical line with tbe tibio-fibnlare. 

 Externally it sends forward a prominent process that, by a trochlear 

 facet at its extremity, articulates with the basal phalanx of the 5th 

 metatarsal. It also articulates with tarsale and metatarsale 4, 

 while in the sole it sends downwards a strong process that serves for 

 muscular and other attachments. This latter is augmented by tbe 

 form assumed by tarsale 4 at its inner side, and it is this common 

 apophysis that gives attachment to some of the short plantar muscles 

 tliat, in my chapter on the myology (given above), may have l)een a 

 little differently described, from the confusing propinquity of tbe 

 ossicles in question. So that a knowledge of this fact will make 

 clear in those premises what might otherwise be considered not an 

 exact description. Professor T. J. Parker, in his studies oi Lacerta 

 viridis, considered tarsale 4 to be the "cuboid" (Zootomy, p. 1.52). 

 Vestiges of a very rudimentary character apjjear to be present in one 

 of my specimens of Ileloderma of a sixth pedal ray, it being in 

 connection with tarsale 5 upon the fibular side of the ankle. 



The metatarsals differ in form but slightly from the metacarpals, 

 and these differences pertain more especially to the proximal extremi- 

 ties, and these have already been pointed out above. Still more in- 

 significant are the differences to be found between the corresponding 

 joints of the digits of manus and pes, and their numerical arrangement 

 is also similar. We saw in the hand that, passing from the radial to 

 the ulnar side, the fingers possessed 2, 3, 4, 5, 3 phalangeal joints, 

 respectively ; now in the foot, in passing from the tibial to the 

 fibular side these numbers exactly correspond, or we find 2, 3, 4, 5, 3 

 phalangeal joints to the toe respectively. 



An excellent diagnostic character twixt pes and manus in this 

 Lizard is to be found in tbe comparative size and form of tbe fifth 

 metatarsal and the fifth metacarpal; in the case of the former it is 

 notably short and small, while in the latter quite tbe reverse of these 

 characters exists, for no especial cnrtailment of its length is to be 

 noticed, and in bulk it rather exceeds any one of the middle three 

 joints of tbe palm. 



