32 2 BOTANICAL GAZETTE [November 



as a usually three-lobed ring just within the inner row of 

 stamens {fig. -?, cp). These lobes ultimately fuse together except 

 the stigmatic tips {fig. i,st)^ and form a rather fleshy ovary wall, 

 with at first a broad, somewhat capitate tip {figs, s^ ^^)- Finally 

 the ovary becomes somewhat barrel-shaped, and when ripe is 

 about 2"^°^ long and 1.5"*"^ in diameter, while the withered stig- 

 mas appear as slight papillae at the upper end {fig. 14, si). 



From an early stage of development there are found in the 

 wall of the ovary numerous scattered oil-secreting cells with 

 nucleated protoplasts {figs. 5^ 11 , oc). In the nearly ripe fruit 

 the carpellary tissue is differentiated into three distinct layers, 

 the outer including the epidermis. The latter is made up of 

 slightly elongated cells with papillose outer walls. At the base 

 of the ovary are found a few scattered, multicellular, pointed 

 hairs similar to those found more abundantly on the base of the 

 spike, and to those shown for Heckeria in fig. 16, The first 

 layer of tissue below the epidermis is of several layers of rather 

 loose parenchyma cells, among which are scattered oil-contain- 

 ing cells {figs. II y 14, 75, <f/'). The second tissue layer is of 

 from five to ten layers of closely packed cells, elongated longi- 

 tudinally to the fruit and without interspersed oil-cells {figs, n, 

 14, 75, cp''). While both these outer layers of the ovary wall 

 are continuous around the whole circumference, the third and 

 innermost layer is discontinuous and confined to the six longi- 

 tudinal grooves in the integument [figs. 14, ij, cp'^^. This layer 

 is made up of somewhat elongated cells WMth trabeculate walls, 

 thickly interspersed with oil-cells. At the border between the 

 second and third layers and opposite the grooves in the integu- 

 ment occur strands of vascular tissue {figs. 11, 14, 15, vb), show- 

 ing from three to eight or ten tracheids in transverse section. 

 These strands continue on into the loose parenchyma of the 

 stylar region where the tracheids become considerably swollen 

 and sometimes detached from each other {figs, //, /-/). 



The ovule, w^hich is terminal on the axis, arises just as the 

 carpel lobes are closing in above. The two integuments appear 

 nearly simultaneously just after the archesporial cell becomes 

 distinguishable {fig. J, tin, otn). The outer integument soon 



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