4o8 BOTANICAL GAZETTE [December 



covered with papillae, specially where the ovary narrows into 

 the style. The papillae in A, Cornuti sometimes develop into 

 long finger-like projections, as observed by Brongniart (2) , 

 although sometimes they form only rounded knobs. Fertiliza- 

 tion causes the rapid development of a dense covering of hairs 

 on the outer surface and sometimes a few on the Inner. Each 

 hair is composed of a single row of cells which arises from an 

 epidermal cell. Another result of fertilization Is the develop- 

 ment of a meristem layer just beneath the inner epidermis. 

 There are signs of this meristem just before fertilization, but it 

 remains only a sign unless fertilization occurs. The result of 

 the activity of this meristem is the rapid thickening of the ovary 



wall, which is accompanied by the formation of dorsiventral 



bundles. Preceding fertilization there is In the carpel only one 

 bundle, centrally located. From this other bundles form later- 

 ally when pod formation begins, but at first all are parallel to 

 the surface of the carpel. The elongated elements which later 

 form the dorsiventral vessels were first noticed when the endo- 

 sperm was in the two to eight-nucleate stage. With the forma- 

 tion of dorsiventral vessels there is a rupture of the parenchyma 

 between them, forming cavities. These are well under way by 

 the time the ^gg undergoes its first division. Further develop- 

 ment of the cavities results in the formation of a pod com- 

 posed of two walls separated by a large space bridged only 

 by bundles with a casing of parenchyma. The cavities extend 

 even into the papillae, which in the mature pod vary from almost 

 nothing to 6"^"^ in length. The dorsal bundles remain closely 

 applied to the dorsal wall of the cavity. The layers of cells 

 next the inner surface become elongated in different directions, 

 forming a firm layer which shows a smooth surface on the side 

 next to the seeds. 



SUMMARY. 



1. The umbels are terminal. 



2. The floral parts appear in centripetal succession. 



3. The members of the same set arise simultaneously, and 

 there is no confluence of primordia. 



4. Glands are present near the axils of the sepals, bracts, and 



I 



