1886.] MR. F. E. BEDDARD ON A LARGE EARTHWORM. 173 
of the penial setae agree with those of A. ungulatus as described by 
Perrier '. 
The ovaries and oviducts I have been unable to find. 
There are two pairs of spermathece situated in segments 8 and 9 ; 
each consists of a spherical thin-walled sac communicating with the 
exterior by a long stout-walled duct which is often curved (see 
figs. 5 and 6, cp); at the upper extremity of the duct, where it unites 
with the sac, it becomes somewhat bulged out on one side, though 
there is hardly so marked a diverticulum as is figured by Perrier 
in A. ungulatus”. 
A very characteristic and remarkable series of structures now 
remain to be described, before concluding the account of the genera- 
tive system. In describing the external characters attention was drawn 
to the modification of the lowermost pair of sete in segment 8; the 
ordinary sete, at any rate on one side of the bedy, have disappeared 
and are replaced by a conspicuous orifice through which protrude 
one or more stout long set, which appear on a naked-eye inspection 
to be very similar to those which project through the male genera- 
tive pores. Fig. 5 of Pl. XIX. represents the internal structures 
which correspond to these peculiarly modified setz; the latter are con- 
tained in a thin-walled transparent sac (s) precisely as are the genital 
sete ; on either side of this sac is a long somewhat sausage-shaped 
glandular body (g.4), which communicates by a slender duct with 
the orifice through which the sete project on to the exterior. The 
presence of these glands renders the whole structure more similar 
still to the male generative pore, except that the “ prostates” are 
paired. These structures were only present on the left side of the 
body in one specimen; in two others they were only developed on 
the right side; in two other specimens both immature, without a 
clitellum, these structures were entirely absent on both sides of 
the body; ina sixth specimen, which was also immature, with the 
elitetum undeveloped and with very minute spermathecee, the bundle 
of modified setze was plainly visible on both sides of the body, but 
without its accessory glands; in the seventh specimen, also imma- 
ture, there was no vestige whatever of these structures ; wherever 
they were absent the setze of the segment were perfectly nurmal. 
I am not aware that any structures of this kind have been 
described in any other species of the genus or in any other Earth- 
worm; they appear to be novel to the group. At the same time 
Perrier figures some structures in A. uxgulatus* which may be 
identical, though the figure and his description are insufficient ; the 
description (p. 92) is as follows :—‘ Les poches copulatrices sont 
situées aux anneaux huit et dix. Chacune est munie d’un lobe 
postérieur, assez petit, et n’a pas d’autre appendice. Dans le neuviéme 
anneau se voient plusieurs sacs glandulaires, séparés par une glande 
multifide”’ The details are evidently different from the structures 
that I have just described, but the similarity of position, between 
the two spermathecze, renders it possible that they are identical ; in 
1 Los, cit. p. 90, pL ii. figs. 21, 22. 
* Loe. cit. pl. ii. fig. 20. 3 Loe, eit. pl. it fig. 18, x. 
