174 MR. F. E. BEDDARD ON A LARGE EARTHWORM. [Mar. 16, 
my spec‘es, however, these structures are in the same segment as that 
which contains the anterior spermatheca and are not in a segment 
intercalated between those which contain the anterior.and posterior 
pairs of spermathece respectively. One of the sete is displayed 
in the drawing (fig. 3), and the lower extremity, more highly 
magnified, in another drawing (fig. 4); the general aspect of these 
setee is very similar to that of the penial setee. The seta is curved 
slightly towards the extremity; its thickness diminishes gradually 
until near to the distal extremity, where it becomes again thicker 
and terminates in a swollen brush-like extremity, the exact shape 
of which can be gathered from an inspection of the figure. The 
distal extremity of the seta is ornamented with delicate transverse 
ridges projecting like the edges of scales and denticulated. 
In Luméricus Hering’ has described, and Vejdovsky * confirmed 
for other species, the modification of certain of the sete in the 
neighbourhood of the genital orifices ; those of the ventral pair in 
“the 10th, 15th, orone of the neighbouring segments, and furthermore 
in the region of the 26th segment and on the clitellum from segment 
31 to segment 38 ”: these sete are more slender than, and double the 
length of, the ordinary sete of the body. In the work referred to 
Vejdovsky goes on to point out that the penial setee in Aeanthodrilus 
and other genera probably correspond to these and differ from the 
genital setze of Chetogaster &c., which are developed during the 
breeding-season in the neighbourhood of the ordinary locomotor 
set, and in addition to them ; the penial setee of Acanthodrilus 
replace the ordinary locomotor sete. Since in Lumbricus the 
modified setze developed in the generative segments are not confined 
to a single segment. or even to the segments bordering upon the 
male generative pores, it is perhaps not surprising to find that in the 
present species of Acanthodrilus there are additional sacs of penial 
setee besides those normally found in the 17th and 19th segments 
of the body. The modified sete of Lumbricus are also furnished 
with a gland which projects into the body-cavity; these are appar- 
ently the capsulogenous glands of D’Udekem and Lankester, which 
are the equivalents of the setigerous glands, being simply enlarged in 
order to assist in the generative function. In Acanthodrilus multi- 
porus I have figured a pair of glands * corresponding to the setz 
which are probably the homologues of these glands, and I imagine 
that in the species of which the present note treats the two large 
glands related to the modified setee of segment 8 are in all proba- 
bility to be referred to the same category and are not special 
structures. 
After the foregoing notes.on the structure of this Earthworm I 
may briefly refer to those points which seem to indicate that it is 
a distinct species differing from both the other two Acanthodrili 
which inhabit New Caledonia. With regard to external characters 
it appears to agree with A. ungulatus in the segments occupied 
1 Zeitschr. f. wiss. Zool. Bd. viii. p. 418. 
? System und Morphologie der Oligochxten (Prag, 1884), pp. 156-7. 
Ann. Se. Nat. 7° série, t. xix. pl. i. fig. 6 4. 
wo 
