72 The Vegetable Individual in its relation to Species. 
male as the fourth ; in Xy/ophylla, the female (on the margins of 
of the female flower, (as in Phyllanthus) as the fifth. In Mo- 
mordica, Eicbalium, Cephalanthera and some other Cucurbitacee, 
the female flower, placed in the axils of the foliaceous leaves of 
the main stem, belongs to the third axis, and the male to the 
fourth ; for the ‘third axis, which here arises from the base of the 
peduncle of the female flower as main axis of the racemose male 
inflorescence, is a superior leaf-shoot. In the other cases,—in which 
the succession of shoots, in order to arrive at the two kinds of 
flowers, separates into two cOordinate lines,—both kinds of flow- 
ers can appear either immediately in the first generation after this 
ration, or, sitice here again preparatory getierations are inter- 
calated, ina later one. Further, the number of the generations 
(axes) in the two lines arising from the division, may be either . 
equal or unequal. A few examples may serve to explain the man- 
ifold cases which thus occur. lu Musa, Myriophyllum and Sag- 
itaria the codrdinate male and female flowers appear in the first 
aie after the separation, and in the whole as a second 
system of axes. Here the female flowers stand in the lower, the — 
male in she upper part of the spicate or racemose inflorescence. 
la eben holds true of Cucurbita and the moneecious Bry- 
;* for here the earlier flowers, which appear in the axils of 
the fi foliaceous leaves, are male; while the later ones which appest 
on the farther pete of the stems are female. A 
below female, in the middle male, and above again ra 4 
flowers, ihodaty en last are dwarfed and sterile. Likewise in 
the first generation after the separation, but in the whole as the 
third system of axes, we find both kinds of flowers in Pachysan- 
dra and Acalypha, and here again, as is usually the case in inde- 
terminate spicate inflorescences of mixed sexes, the female flower 
is in the lower, the male in the upper part of the inflorescence. 
he same obtains in moneecious Palms with axillary spadices ; 
thongh here the flowers appear in ramified spikes from the fourth 
system of axes. When the flowers make their appearance in the 
second generation after the division, they cannot easily be united 
in the same st age and special male and female inflores- 
cences will ar Thus, e.g., in Plutanus, Liquidambar and 
ecanisal in in which the female inflorescences occur on the 
lower part of the main shoot, and the male in the upper; like- 
wise in Quercus and Fagus, though here, vice versa, the male 
inflorescences are the lower, and the female the upper. eee 
if the division of the succession of shoots is an unequal o1 ein 
* Bryonia has apparentl: investigation 
that ny xy do not spr sti pe incodenie ong g so te the pee leaf, but ct (ee 
=) out 0 of the peduncle of a aber racoat directly in the 
axil of the leaf which exactly eeceapaade ts vee lowes 
+ The inflo —— well a6 Gad th Cait 
