120 "White. 



4. The race (300 — 309) is such a homozyg-ous pure line because 

 it has been propagated from seed obtained by selfing a single (original) 

 plant, which is believed to have arisen or mutated by a single factor A 

 from such a homozygous pure line. 



5. The factor A of this race, in expressing itself somatically, 

 when surrounded by its original gene complex (402) affects a large 

 number of important plant organs. Prominent among these may be 

 mentioned the stem, the leaf number and arrangement (phyllotaxy), the 

 inflorescence and the flowers. The nature of this expression is held 

 to be ontogenetically progressive, as the seedlings and the early juvenile 

 stages do not show any distinguishing fasciata features. As the plants 

 of this race progress toward maturity, the factor more and more 

 implants its distinctive morphological characteristics upon the various 

 organs, so that those wliich develop last, exhibit the greatest alterations. 

 Hence, the last flower whorl laid down in ontogenetic development is 

 the most altered from its normal expression by the factor A. 



6. The hereditary nature of the fasciated condition has been 

 tested by breeding large numbers of progeny from the seeds of a single 

 selfed plant. In all cases, the character is constant in the sense that 

 its extreme fluctuations do not approach the normal (402) condition 

 near enough so as to call forth any question as to which is which when 

 the two are grown together. In other words, all the progeny of selfed 

 plants of this race express the character fasciation, as described under 

 "materials" to some degree. Its fluctuation is largely "inherent" and 

 not the result of the "external" environments under which the ex- 

 perimental cultures were grown. As no "atavists" appeared, the character 

 is not "eversporting" in the de Vriesian sense, 



7. The repetition in the number of organs, such as leaves, sepals, 

 petals, stamens and ovary -locules is not a duplication of whorls 

 (pleiotaxy) or of practically whole organisms, as the theories of some 

 anatomists would seem to imply, and as de Vries suggests in explanation 

 of his data on Geranium molle fasciatum. No evidence of congenital 

 mechanical fusions is given by cross-sections of the mature stem. The 

 different whorls in the flower appear to vary somewhat independently 

 of each other, as the correlation in number of parts between those of 

 a single flower is far from perfect, though probably exceeding 50 per 

 cent. The progressive expression of the factor in its ontogenetic 

 development may entirely account for this. 



