Genetical Studies on Oenothera. V. 191 



sepals much less heavy, resembling that of sepals very heavy resembling that of 



of biennis. franeiscana. 



Petals. 21— 2-4 cm. long. Petals. 2-3— 2-5 cm long. 



Stigma lobes. 6 — 7 mm. long, varying Stigma lobes. 6 — 7 mm. long, varying 



from slightly above to slightly below from slightly above to slightly below 



the tips of the anthers, longer than those the tips of the anthers, longer than those 



of either parent. of either parent. 



Capsules. 2 — 2'8 cm. long, shorter and less Capules. 2"2 — .3 cm. long, longer and more 



heavily pubescent, resembling biennis. heavily pubescent, resembling /Vanascana. 



Papillae red. Papillae red. 



It is clear that the reciprocal crosses of biennis and franeiscana 

 present a situation similar to that described for the reciprocals of 

 biennis and muricata. The reciprocal crosses exliibit striking contrast- 

 ing differences and in certain characters strongly resemble the pollen 

 parents. The points of greatest contrast are (1) in the leaf forms of 

 the young and old rosettes and in the degree of crinkling, (2) in the 

 degree of pubescence on the stems, sepals, and capsules, (3) in the size 

 and habit of the mature plants, and in the forms of the foliage and 

 bracts, (4) in the character of the sepal tips and in the form of the 

 bud cone, (5) in the length of the capsules. The flower structure of 

 the hybrids, with a somewhat wide range of variation, does not present 

 very striking differences in the size of the petals or in the position of 

 the stigma relative to the tips of the anthers although the parents in 

 these characters offer sharp contrasts. 



The patroclinous tendencies are expressed most strongly in the 

 contrasting characters listed in the paragraph above. The hybrids 

 proved to be matroclinous in the time of the appearance of the side 

 branches from the rosette relative to the growth of the central shoot. 

 The influence of the maternal parent upon the hybrids was apparent 

 even when characters were most strongly patroclinous and I found no 

 certain evidence that any parental character is ever exactly duplicated 

 in the hybrid except that of red coloration considered below. 



In one character there was presented a form of relative dominance 

 of franeiscana over biennis. The red coloration of the stem papillae 

 and of the sepals conspicuous in franeiscana and absent in biennis, 

 was exhibited by both hybrids. Thus franeiscana X biennis was ma- 

 troclinous with respect to these color characters and biennis X franeis- 

 cana was patroclinous. The depth of red in the sepals of the hybrids 

 varied somewhat widely, in some plants being much lighter than that 

 of the parent franeiscana and in other plants being fully as dark. The 

 relative numbers of papillae over the stems of the hybrids followed 



