Genetical Stiulies on Oenothera. V. 193 



The race grandiffora U is representative of the hroader-leaved forms of 

 grandiffora. 



It is only necessary at this time to g-ive such characters of gran- 

 difiora I) as offer the most striidng points of contrast to those of the 

 Dutch biennis, which characters may l)e compared w'ith the description 

 of the latter species to be found on pages 174 and 175 of this paper. 



Mature rosettes. 2 — 2'ö dm. broad. A loose cluster of broadly-elliptical leaves, irregu- 

 larly and sometimes deeply toothed at the base, with reddish blotches and but 

 slightly crinkled (see Davis '11, Fig. (i). The rosettes are transitory sending up 

 very quickly the central shout and consequently under favorable conditions 

 strictly annual. 



Mature plants. 1'5 — 2 m. high. A strong central shoot frequently with long branches 

 from the base of the plant, generally profusely branched above (see Davis '11, 

 Fig. 7). Stems reddish, green only at their tips, papillae following the color of 

 the stem. 



Foliage. Leaves below ovate or broadly-elliptical, scarcely half the length of the 

 leaves of biennis. 



Inflorescence. Bracts '/i — V2 length of the mature buds, broadly-lanceolate, persistent 

 so that the fruiting branches remain leafy (see Davis '11, Fig. 8); those of 

 biennis are likely to lose their bracts. Buds less closely set in the spike than 

 in biennis. 



Mature buds. 9 — 10 cm. long. Cone 4 — b cm. long, scarcely angled. Sepals marked 

 with reddish spots and blotches. Sepal tips 9 — 12 mm. long, very slender (see 

 Davis '11, Fig. 4). 



Petals. 3'3 — 3'5 cm. long (see Davis '11, Fig. 8). Flowering period late in the season 

 {biennis flowers early). 



Stigma lobes. 6 — 7 mm. long, generally 2 — '^ mm. above the tips of the anthers; those 

 of biennis are usually about .3 mm. below. 



Capsules. 2 5 — 3 cm. long. 



From 261 seeds of the cross grand! flora X biennis 170 seedlings 

 were obtained from seed pans kept for 5 weeks; 131 plants (culture 

 12"48) were brought to maturity. In the reciprocal cross, biennis X grandi- 

 flora, 174 seeds gave after 5 weeks in the seed pans 120 seedlings of 

 which 112 plants matured (culture 12"49). The two cultures presented 

 the following contrasting characters: 



12'48, yrandiflora X biennis. 12"49, biennis ',\ grandiflora. 



Mature rosettes. Leaves broader and with Mature rosettes. Leaves narrower and with 

 shorter petioles, and consequently some- longer petioles, and consequently some- 

 what more closely clustered (a patro- what more loosely clustered (a patro- 

 clinous tendency). Clear evidence of a clinous tendency). Clear evidence of a 

 blend of the parental characters. Some blend of the parental characters. Some 

 rosettes heavily blotched with red, some rosettes heavily blotched with red, some 

 almost clear green. Patroclinous in the almost clear green. Patroclinous in 

 delayed appearance of the central shoot. quickly sending up the central shoot. 



