Sex-limited inheritance in Lychnis dioica L. 275 



X£i^.Z6i^ = heterozygous broad-leafed females, 

 XhF .XhF ^ ]io\\\ozygo\x?, narrow-leafed females, 

 XBF.Xhf = heterozygous broad-leafed males, 

 XbF.Xbf = homozygous narrow-leafed males. 



It will be noted that tliis expectation differs from that for the Fj 

 in only one point, namely, in the appearance of homozygous narrow- 

 leafed females instead of homozygous broad-leafed females. I confidently 

 predicted the fulfilment of these expectations and laid plans for the 

 extensive use of narrow-leafed females in subsequent breeding; imagine 

 my astonishment at the results presented in Table I. 



Table I. 

 Genetic formula: XBF.XfFX XbF.Xff. 



It is seen that the two expected rosette-tyjjes, broad and narrow, 

 occurred in each family, in ratios not more divei-gent from the expected 

 1 : 1, than might well be accounted for by the obviously less vigorous 

 constitution of the angustifolia-iovm, but in the entire series of families, 

 aggregating 1097 individuals, there were only 2 females! No adequate 

 explanation can be offered at the present time for this remarkable result. 

 It must be made the subject of further experimental inquiry. That there 



