222 ; The Botanical Gazette. [June, q 
of cells persist. As development goes on, the outer layer 
becomes inconspicuous on account of the obliteration of the 
cell lumina and the almost hyaline appearance of the walls, 
The inner layer suffers less compression usually on account 
of the thicker walls, but often treatment with potassic hydrate 
is necessary to bring out the elements. With the exception 
of the region near the fibro-vascular bundle and the epidermis, 
the nucellus is eventually all resorbed. The epidermis per- 
sists in a compressed condition. 5 
At maturity, the ovary wall outside of the chlorophyll 
layer consists of two or three rows of cells with strongly 
thickened walls and narrowed lumina, fig. 13. The space formerly 
occupied by the absorbed portion of the wall has been filled 
by the outward movement of the inner growing parts thus 
bringing the chlorophyll layer in contact with the remaining 
cells of the ovary wall. The remaining cells of the inner 
epidermis, when present, and, in their absence, the former 
chlorophyll-bearing cells, are soldered to the hyaline outer 
layer of the inner integument and sometimes also exteriorly to 
the adjacent cells of the wall. Of the existence of this union, 
I think there can be no doubt. The inner integument, said 
by Jumelle to be absent, was present in every specimen I have 
examined and is plainly distinguishable under moderate 
magnification in specimens that have been treated with 
potassic hydrate. The hyaline outer layer is not ¢asi 
demonstrated, but the thicker walled inner layer is readily 
seen, fig. 12. : all 
In ‘‘red” wheats, particularly, this is true, since in the ¢ | 
Cavities of this layer is deposited a quantity of ae se 
coloring material that catches the eye at once. The pe 
€nce in color between the so-called red and white w 
depends on the presence or absence of this coloring geo 
the inner integument. As far as I have been able ee 
tain, this layer of cells is always present in the white W 
but is less conspicuous than in the red. ian ak 
Thinking that possibly a difference between America® 
Ist 
European varieties might account for the difie al 
