26 BOTANICAL GAZETTE [juLY 
nuclei of the sheath cells of the archegonium, and thinks he has 
traced their entrance through protoplasmic connections in several 
species of Pinus and in Adves sibirica. From his work on Dam- 
mara and Cephalotaxus he is inclined to consider their origin as 
the same in these cases also. While I have not been able to 
establish the existence of protoplasmic connections between egg 
and sheath cells, the appearance in fig. 83 strongly suggests that 
such connections exist, and the presence of pits in the wall of 
the archegonium would also imply their occurrence. 
The two denser areas already noticed in the very young 
archegonium (fig. 62) have reached in fig. 63 a size and condi- 
tion retained until the initial changes which bring on the division 
into the central canal and egg nuclei. These areas are of dense 
fibrous material, and are by far the most striking features in the 
archegonium of Taxodium. They stain much more deeply with 
orange G than does the surrounding cytoplasm, and from them 
fibers radiate to the surface of the cell. It will be noticed that 
the denser part is at the periphery of the mass, but the inner 
part is also denser than the ordinary cytoplasm of the cell, and the 
whole is composed of a complex of granular fibers. The upper 
of these masses isthe smaller and lies very near the nucleus. 
Fibers can be traced passing from the cen‘ral mass around the 
nuclear wall, and they seem to be a continuation of the wall itself. 
That these bodies are of the same nature as the so-called kino- 
plasmic material, generally most conspicuous at the time of 
nuclear division, is evident. They show the same structure as 
the much less developed kinoplasmic areas already mentioned 
in the megaspore. Such bodies have not heretofore been 
described as occurring in such perfection in any case with which 
Iam acquainted. Kinoplasmic areas have been mentioned near 
the nucleus at the time of its division in the archegonium of 
Tsuga canadensis by Murrill (’00), and Ikeno (’98) has figured 
such areas under the central cell nucleus of Cycas. Chamberlain 
(’99) gives one case where there are two such bodies near the 
egg nucleus in Pinus Laricio, and Blackman (’98) describes fibers 
of this nature radiating from the egg nucleus before fertilization. 
In all these cases, however, the fibrous material is not nearly so 
