130 BOTANICAL GAZETTE [AUGUST 
Such a change will require us to discard the name Taxodieae 
and combine the remaining genera of the family into some ten- 
tative group of another name until further study shall make their 
position clear. 
SUMMARY. 
The staminate cones begin to develop in September or Octo- 
ber, and by winter the pollen mother-cells are formed. In spring 
starch is removed from the cells of the sporophyll and stored in 
the mother-cells, where it remains through their divisions and 
disappears in the pollen grain as the exine is being formed. The 
exposed wall of the microsporophyll is but two layers thick. 
The reducing divisions in the pollen mother-cells resemble 
those in the Larix and the reduced number of chromosomes is 
probably twelve. There is a fairly well developed resting stage 
after the first division in the mother-cells. 
About ten days after the reducing division a division of the 
pollen grain occurs which separates at once the generative cell 
from the tube cell. No sterile prothallial cells are formed. 
From two to three weeks after pollination, when the pollen 
tube has grown some distance, the generative cell divides into 
the central cell and stalk cell, and these move down toward the 
tube nucleus. The pollen tube reaches the prothallium earlier 
than in any case previously described, sometimes even before 
the formation of a cellular tissue in the latter. 
The arrangement of the nuclei in the pollen tube is the same 
as in other conifers. 
The central cell has a distinct Hautschicht of its own and 
resembles in outline that of Taxus and the Cupresseae. It 
divides simultaneously with the division in the central cell of the 
archegonium, and the two sperm cells thus formed move apart 
slightly. They are furnished with a dense layer of starch around 
the nucleus, a peripheral finely granular layer often containing 
globules of plastic material, and a Hautschicht. Its nucleus is 
densely filled with granular material and has a coarse chromatin 
recticulum and a nucleolus. 
The ovulate cones also begin their development in early fall 
and continue growing slowly, as the weather permits, through 
