336 BOTANICAL GAZETTE [NOVEMBER 
difficulty of finding a perfect underleaf. An examination of the 
specimens of A. mucosa which were distributed in Hepaticae Spruce- 
anae shows that the secretion of slime is performed by club- 
shaped papillae, similar in all respects to those described for 
Odontoschisma, and that it is not necessarily accompanied by 
the destruction of cells. The only difference between the genera 
in this respect is a difference of degree, the underleaves of 
Anomoclada being larger and the papillae more numerous than 
in Odontoschisma. They arise not only from the margin of an 
underleaf, but also from the postical surface near the margin, and 
sometimes a cell is directly transformed into a papilla. The basal 
cells of the underleaves acquire thickened walls with distinct tri- 
gones, and persist indefinitely, even after the papillae have lost 
their protoplasm and withered away. According to Spruce, 
the underleaves are ‘late ovata in acumen subulamve brevem 
producta . . . . superiora vix unquam perfecta, sed e margine 
apiceque plus minus dissolutis, nunc irregulariter bifida, nunc 
quadrifida vel digitatim multifida.” As a matter of fact, the 
underleaves are variously divided, even in the vicinity of the 
growing point, and this division is in no sense due to the devel- 
opment of slime-secreting papillae. There seems to be no good 
reason, therefore, for considering that they are primarily undi- 
vided. 
The strong resemblance between the underleaves of Odonto- 
schisma and Anomoclada indicates a close relationship between 
the genera. They resemble each other further in their prostrate 
stems with postical flagella, in their succubous undivided leaves, 
and in their thick-walled leaf-cells with distinct trigones. Even 
closer to Anomoclada than any of the species which have yet 
been noted, is Odontoschisma Portoricense,a West Indian species 
(figs. 65-74). At first sight this looks precisely like a poorly 
developed form of A. mucosa, largely from the fact that its 
leaves are commonly crispate near the postical base and slightly 
convex—peculiarities which none of the more northern forms of 
the genus show. In O. Portoricense the underleaves (fig. 70) are 
covered over with slime-papillae, and the perichaetial bracts are 
often slightly connate (fig. 77), the latter being a character 
