Occurrence of different systems of gametic reduplioation. 29 



The results of Oenothera were cytoloufically exi)laine(l l)y (iOLD- 

 SCHMIDt') as due to a merogony, wliile Rennek comes to an oi)i)osite 

 conclusion-). The question remains still unsettled. 



Gregory's case was criticised by Trow ■') who maintains that tht- 

 case might have been due to 2:1:1:2 coupling, the system which 

 he first found in Senccio vulgaris, occurring alike l)otii in male and 

 female gametes. 



Saunders' conclusions were also discussed l)y Goldschmidt ^) 

 in supposing the case as an example of sex -limited inheritance. The 

 criticism was replied quite recently by the original authoress ■'). At 

 any rate, the difficulty for a critical proof of her hypothesis rests, 

 perhaps, upon the fact that the reproductive organs of the female (in 

 Petunia), or of both sexes (in Maffhiola) of the recessive (doul)le- 

 flowering) plants are totally sterile. 



The essential feature by which my present results differ from those 

 of DE Vries and Sauxders is that the dissimilarity of gametic distribut- 

 ion in both sexes is quantitative in its nature, because the cases may 

 be considered as the reduplication which is low i)artial in the male is 

 rendered, from unknown cause, very intense in the female, while the 

 difference is assumed to be qualitative in the cited cases of the previous 

 authors. 



It should be noticed that in the present experiments with silkworms, 

 though peculiar in the mode of inheritance, we are by no means deal- 

 ing with the characters which are sex -limited. Fis ex A Abb 9 X 

 aaBB c? are of the same kind as those ex aaBB 9 X AAbb cf, and 

 the result of AABB 9 X aabh cT is always similar to that of aabh 

 9 X AABB cf. In a sex -limited inheritance, on the contrary, the 

 reciprocal crosses will give, as a rule, Fi series decidedly differed from 

 each other. 



') Die Merogonie der Oeno(Afra-Bastarde und die doppeltreziproken Bastarde von 

 DE Veies. Arch. f. Zellforsch., Bd. 9, 1912. Ref. in Zeitschr. f. ind. Abst. u. 

 Vererb. Bd. IX, 191.% pp. 135 — 1,36. 



') Über die augebliche Merogonie der Ocno/Acra- Bastarde. Ber. d. deutsch, 

 bot. Gesell., Bd. XXXI, 191.% pp. ;^;34 -3.3.^. 



•) On the Inheritance of Certain Characters in the Common Groundsel. Journ. 

 Gen. Vol. 2, No. 3, 1912, p. 272. 



*) Der Vererbungsmodus der gefüllten Levkojenrassen als Fall geschlechtsbegrenzter 

 Vererbung? Zeitschr. f. ind. Abst. u. Vererb. Bd. 10, Ht. I u. 2, 1913, pp. 74— 9b. 



') On the Mode of Inheritance of certain Characters in Double-throwing Stocks, 

 Zeitschr. f. ind. Abst. u. Vererb. Bd. 10, Ht. 4, 191.3, pp. 297— 310. 



