2i6 Gates. 



In fig. I is shown a group of seedlings of grandijlora of varying 

 age as they appeared in my cultures from seeds sown in September 

 1909. In fig. 2 appears a later stage in rosette development, the two 

 smaller ones showing certain marked differences from the type. It 

 will be observed that indistinct (pale red) blotches appear on the older 

 rosettes. There is obviously a considerable range of variability in 

 these plants, but all of them which fully developed belonged to one well- 

 marked type. Under certain conditions of culture the rosette shown 

 in fig. 2 is followed by a stage in which a new and very characteristic 

 type of leaf with deep basal lobes appears (see Gates 1913 a, PI. IV, 

 figs. 46, 47), but this is always suppressed in all forms of 0. grandi- 

 jlora and their hybrids, when grown as annuals. 



Wlaen set out-of-doors a stem very soon shoots up and the rosette 

 disappears. The plants soon come into flower, and the result is an 

 adult plant having, in the climate of St. Louis, Mo., the appearance of 

 fig. 3. The plants were tall but with slender stems, and the lowermost 

 branches were some distance above the ground, giving the plant, 

 which frequently developes small secondary branches, a bushy aspect. 

 Last year I grew in the English climate a culture of two hundred and 

 twenty-one plants from the same source and found several striking 

 changes in their developmental physiology. As usual, they formed 

 a rather evanescent rosette and began stem fonnation long before 

 the other Oenothera races, but though they developed luxuriantly 

 in a vegetative way they failed entirely to produce flowers, only two 

 plants bearing a few buds and flowers at the end of the season. In this 

 behaviour they agreed with a race of 0. Tracy i Bartlett from the 

 same climate — ■ Birmingham, Alabama. Some of the plants branched 

 from the base, giving the appearance shown in fig. 4. I should mention 

 that this culture produced one dwarf, presumably a mutation. This 

 plant was unbranched and only two feet high, but otherwise resembled 

 grandijlora. 



Fig. 5 is a row of buds from my 191 1 culture of grandijlora, two- 

 thirds natural size, showing their smooth, rounded, slender character 

 as contrasted with those of ruhricalyx (fig. 9), and the range of the red 

 colovu" pattern on their sepals. 



The main distinctive features of grandijlora may be summarized 

 as follows (cf. table I): The leaves are thin and light green, the rosette 

 leaves bearing characteristic red blotches. The rosettes are evanescent, 

 the plants "early" in development, the stem-leaves always pointed 

 at both ends. This shows particularh' well in fig. 4. The bud characters 



